Transpiration in the context of Water circulation

The water cycle (or hydrologic cycle or hydrological cycle) is a biogeochemical cycle that involves the continuous movement of water on, above and below the surface of the Earth across different reservoirs. The mass of water on Earth remains fairly constant over time. However, the partitioning of the water into the major reservoirs of ice, fresh water, salt water and atmospheric water is variable and depends on climatic variables. The water moves from one reservoir to another, such as from river to ocean, or from the ocean to the atmosphere due to a variety of physical and chemical processes. The processes that drive these movements, or fluxes, are evaporation, transpiration, condensation, precipitation, sublimation, infiltration, surface runoff, and subsurface flow. In doing so, the water goes through different phases: liquid, solid (ice) and vapor. The ocean plays a key role in the water cycle as it is the source of 86% of global evaporation.

The water cycle is driven by energy exchanges in the form of heat transfers between different phases. The energy released or absorbed during a phase change can result in temperature changes. Heat is absorbed as water transitions from the liquid to the vapor phase through evaporation. This heat is also known as the latent heat of vaporization. Conversely, when water condenses or melts from solid ice it releases energy and heat. On a global scale, water plays a critical role in transferring heat from the tropics to the poles via ocean circulation.

In botany, a stoma (pl.: stomata, from Greek στόμα, "mouth"), also called a stomate (pl.: stomates), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange between the internal air spaces of the leaf and the atmosphere. The pore is bordered by a pair of specialized parenchyma cells known as guard cells that regulate the size of the stomatal opening.

The term is usually used collectively to refer to the entire stomatal complex, consisting of the paired guard cells and the pore itself, which is referred to as the stomatal aperture. Air, containing oxygen, which is used in respiration, and carbon dioxide, which is used in photosynthesis, passes through stomata by gaseous diffusion. Water vapour diffuses through the stomata into the atmosphere as part of a process called transpiration.

Soil moisture is the water content of the soil. It can be expressed in terms of volume or weight. Soil moisture measurement can be based on in situ probes (e.g., capacitance probes, neutron probes) or remote sensing methods.

Water that enters a field is removed from it by runoff, drainage, evaporation or transpiration. Runoff is the water that flows on the surface to the edge of the field; drainage is the water that flows through the soil downward or toward the edge of the field underground; evaporative water loss from a field is that part of the water that evaporates into the atmosphere directly from the field's surface; transpiration is the loss of water from the field by its evaporation from the plant itself.

Evapotranspiration (ET) refers to the combined processes which move water from the Earth's surface (open water and ice surfaces, bare soil and vegetation) into the atmosphere. It covers both water evaporation (movement of water to the air directly from soil, canopies, and water bodies) and transpiration (evaporation that occurs through the stomata, or openings, in plant leaves). Evapotranspiration is an important part of the local water cycle and climate, and measurement of it plays a key role in water resource management and agricultural irrigation.

Plant physiology is a subdiscipline of botany concerned with the functioning, or physiology, of plants.

Plant physiologists study fundamental processes of plants, such as photosynthesis, respiration, plant nutrition, plant hormone functions, tropisms, nastic movements, photoperiodism, photomorphogenesis, circadian rhythms, environmental stress physiology, seed germination, dormancy and stomata function and transpiration. Plant physiology interacts with the fields of plant morphology (structure of plants), plant ecology (interactions with the environment), phytochemistry (biochemistry of plants), cell biology, genetics, biophysics and molecular biology.

Drylands are defined by a scarcity of water. Drylands are zones where precipitation is balanced by evaporation from surfaces and by transpiration by plants (evapotranspiration). The United Nations Environment Program defines drylands as tropical and temperate areas with an aridity index of less than 0.65. Drylands can be classified into four sub-types:

• Dry sub-humid lands
• Semi-arid lands
• Arid lands
• Hyper-arid lands

Some authorities regard hyper-arid lands as deserts (United Nations Convention to Combat Desertification) although a number of the world's deserts include both hyper-arid and arid climate zones. The UNCCD excludes hyper-arid zones from its definition of drylands.

A tracheid is a long and tapered lignified cell in the xylem of vascular plants. It is a type of conductive cell called a tracheary element. Angiosperms also use another type of conductive cell, called vessel elements, to transport water through the xylem. The main functions of tracheid cells are to transport water and inorganic salts, and to provide structural support for trees. There are often pits on the cell walls of tracheids, which allows for water flow between cells. Tracheids are dead at functional maturity and do not have a protoplast. The wood (softwood) of gymnosperms such as pines and other conifers is mainly composed of tracheids. Tracheids are also the main conductive cells in the primary xylem of ferns.

The tracheid was first named by the German botanist Carl Gustav Sanio in 1863, from the German Tracheide.

A cactus (pl.: cacti, cactuses, or less commonly, cactus) is a member of the plant family Cactaceae (/kækˈteɪsi.iː, -ˌaɪ/), a family of the order Caryophyllales comprising about 127 genera with some 1,750 known species. The word cactus derives, through Latin, from the Ancient Greek word κάκτος (káktos), a name originally used by Theophrastus for a spiny plant whose identity is now not certain. Cacti occur in a wide range of shapes and sizes. They are native to the Americas, ranging from Patagonia in the south to parts of western Canada in the north, with the exception of Rhipsalis baccifera, which is also found in Africa and Sri Lanka. Cacti are adapted to live in very dry environments, including the Atacama Desert, one of the driest places on Earth. Because of this, cacti show many adaptations to conserve water. For example, almost all cacti are succulents, meaning they have thickened, fleshy parts adapted to store water. Unlike many other succulents, the stem is the only part of most cacti where this vital process takes place. Most species of cacti have lost true leaves, retaining only spines, which are highly modified leaves. As well as defending against herbivores, spines help prevent water loss by reducing air flow close to the cactus and providing some shade. In the absence of true leaves, cacti's enlarged stems carry out photosynthesis.

Cactus spines are produced from specialized structures called areoles, a kind of highly reduced branch. Areoles are an identifying feature of cacti. As well as spines, areoles give rise to flowers, which are usually tubular and multipetaled. Many cacti have short growing seasons and long dormancies and are able to react quickly to any rainfall, helped by an extensive but relatively shallow root system that quickly absorbs any water reaching the ground surface. Cactus stems are often ribbed or fluted with a number of ribs which corresponds to a number in the Fibonacci numbers (2, 3, 5, 8, 13, 21, 34 etc.). This allows them to expand and contract easily for quick water absorption after rain, followed by retention over long drought periods. Like other succulent plants, most cacti employ a special mechanism called "crassulacean acid metabolism" (CAM) as part of photosynthesis. Transpiration, during which carbon dioxide enters the plant and water escapes, does not take place during the day at the same time as photosynthesis, but instead occurs at night. The plant stores the carbon dioxide it takes in as malic acid, retaining it until daylight returns, and only then using it in photosynthesis. Because transpiration takes place during the cooler, more humid night hours, water loss is significantly reduced.

Water-use efficiency (WUE) refers to the ratio of plant biomass to water lost by transpiration, can be defined either at the leaf, at the whole plant or a population/stand/field level:

• leaf level : photosynthetic water-use efficiency (also called instantaneous water-use efficiency WUE_inst), which is defined as the ratio of the rate of net CO₂ carbon assimilation (photosynthesis) to the rate of transpiration or stomatal conductance, then called intrinsic water-use efficiency (iWUE or W_i)
• plant level : water-use efficiency of productivity (also called integrated water-use efficiency or transpiration efficiency,TE), which is typically defined as the ratio of dry biomass produced to the rate of transpiration.
• field level : based on measurements of CO₂ and water fluxes over a field of a crop or a forest, using the eddy covariance technique

Research to improve the water-use efficiency of crop plants has been ongoing from the early 20th century, however with difficulties to actually achieve crops with increased water-use efficiency.

In viticulture, the canopy of a grapevine includes the parts of the vine visible aboveground - the trunk, cordon, stems, leaves, flowers, and fruit. The canopy plays a key role in light energy capture via photosynthesis, water use as regulated by transpiration, and microclimate of ripening grapes. Canopy management is an important aspect of viticulture due to its effect on grape yields, quality, vigor, and the prevention of grape diseases. Various viticulture problems, such as uneven grape ripening, sunburn, and frost damage, can be addressed by skillful canopy management. In addition to pruning and leaf trim, the canopy is often trained on trellis systems to guide its growth and assist in access for ongoing management and harvest.

Polytrichaceae is a common family of mosses. Members of this family tend to be larger than other mosses, with the larger species occurring in particularly moist habitats. The leaves have specialized sheaths at the base and a midrib that bears photosynthetic lamellae on the upper surface. These mosses are capable of sustaining high rates of photosynthesis in the presence of ample light and moisture. Unlike all other mosses, the hydroid-based vascular system of these mosses is continuous from stem to leaf and can extract water from the soil through transpiration. Species in this group are dioicous, though some are monoicous. In most species, the sporophytes are relatively large, the setae are rigid, and the calyptrae are hairy. Most species have nematodontous peristomes with 32–64 teeth in their sporangium; some early-diverging genera instead have a stopper mechanism, which consists of the apical section of the columella, that seals the mouth of the capsule shut prior to dehiscence.

Ecohydrology (from Greek οἶκος, oikos, "house(hold)"; ὕδωρ, hydōr, "water"; and -λογία, -logia) is an interdisciplinary scientific field studying the interactions between water and ecological systems. It is considered a sub discipline of hydrology, with an ecological focus. These interactions may take place within water bodies, such as rivers and lakes, or on land, in forests, deserts, and other terrestrial ecosystems. Areas of research in ecohydrology include transpiration and plant water use, adaption of organisms to their water environment, influence of vegetation and benthic plants on stream flow and function, and feedbacks between ecological processes, the soil carbon sponge and the hydrological cycle.

In plants, the transpiration stream is the uninterrupted stream of water and solutes which is taken up by the roots and transported via the xylem to the leaves where it evaporates into the air/apoplast-interface of the substomatal cavity. It is driven by capillary action and in some plants by root pressure. The main driving factor is the difference in water potential between the soil and the substomatal cavity caused by transpiration.