Transpiration is the process of water movement through a plant and its evaporation from aerial parts, such as leaves, stems and flowers. It is a passive process that requires no energy expense by the plant. Transpiration also cools plants, changes osmotic pressure of cells, and enables mass flow of mineral nutrients. When water uptake by the roots is less than the water lost to the atmosphere by evaporation, plants close small pores called stomata to decrease water loss, which slows down nutrient uptake and decreases CO2 absorption from the atmosphere limiting metabolic processes, photosynthesis, and growth.
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👉 Transpiration in the context of Stomata
In botany, a stoma (pl.: stomata, from Greek στόμα, "mouth"), also called a stomate (pl.: stomates), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange between the internal air spaces of the leaf and the atmosphere. The pore is bordered by a pair of specialized parenchyma cells known as guard cells that regulate the size of the stomatal opening.
The term is usually used collectively to refer to the entire stomatal complex, consisting of the paired guard cells and the pore itself, which is referred to as the stomatal aperture. Air, containing oxygen, which is used in respiration, and carbon dioxide, which is used in photosynthesis, passes through stomata by gaseous diffusion. Water vapour diffuses through the stomata into the atmosphere as part of a process called transpiration.
In this Dossier
- ⭐ Core Definition: Transpiration
- 👉 Transpiration in the context of Stomata
- Transpiration in the context of Water cycle
- Transpiration in the context of Soil moisture
- Transpiration in the context of Evapotranspiration
- Transpiration in the context of Plant physiology
- Transpiration in the context of Drylands
Transpiration in the context of Water cycle
The water cycle (or hydrologic cycle or hydrological cycle) is a biogeochemical cycle that involves the continuous movement of water on, above and below the surface of the Earth across different reservoirs. The mass of water on Earth remains fairly constant over time. However, the partitioning of the water into the major reservoirs of ice, fresh water, salt water and atmospheric water is variable and depends on climatic variables. The water moves from one reservoir to another, such as from river to ocean, or from the ocean to the atmosphere due to a variety of physical and chemical processes. The processes that drive these movements, or fluxes, are evaporation, transpiration, condensation, precipitation, sublimation, infiltration, surface runoff, and subsurface flow. In doing so, the water goes through different phases: liquid, solid (ice) and vapor. The ocean plays a key role in the water cycle as it is the source of 86% of global evaporation.
The water cycle is driven by energy exchanges in the form of heat transfers between different phases. The energy released or absorbed during a phase change can result in temperature changes. Heat is absorbed as water transitions from the liquid to the vapor phase through evaporation. This heat is also known as the latent heat of vaporization. Conversely, when water condenses or melts from solid ice it releases energy and heat. On a global scale, water plays a critical role in transferring heat from the tropics to the poles via ocean circulation.
Transpiration in the context of Soil moisture
Soil moisture is the water content of the soil. It can be expressed in terms of volume or weight. Soil moisture measurement can be based on in situ probes (e.g., capacitance probes, neutron probes) or remote sensing methods.
Water that enters a field is removed from it by runoff, drainage, evaporation or transpiration. Runoff is the water that flows on the surface to the edge of the field; drainage is the water that flows through the soil downward or toward the edge of the field underground; evaporative water loss from a field is that part of the water that evaporates into the atmosphere directly from the field's surface; transpiration is the loss of water from the field by its evaporation from the plant itself.
Transpiration in the context of Evapotranspiration
Evapotranspiration (ET) refers to the combined processes which move water from the Earth's surface (open water and ice surfaces, bare soil and vegetation) into the atmosphere. It covers both water evaporation (movement of water to the air directly from soil, canopies, and water bodies) and transpiration (evaporation that occurs through the stomata, or openings, in plant leaves). Evapotranspiration is an important part of the local water cycle and climate, and measurement of it plays a key role in water resource management and agricultural irrigation.
Transpiration in the context of Plant physiology
Plant physiology is a subdiscipline of botany concerned with the functioning, or physiology, of plants.
Plant physiologists study fundamental processes of plants, such as photosynthesis, respiration, plant nutrition, plant hormone functions, tropisms, nastic movements, photoperiodism, photomorphogenesis, circadian rhythms, environmental stress physiology, seed germination, dormancy and stomata function and transpiration. Plant physiology interacts with the fields of plant morphology (structure of plants), plant ecology (interactions with the environment), phytochemistry (biochemistry of plants), cell biology, genetics, biophysics and molecular biology.
Transpiration in the context of Drylands
Drylands are defined by a scarcity of water. Drylands are zones where precipitation is balanced by evaporation from surfaces and by transpiration by plants (evapotranspiration). The United Nations Environment Program defines drylands as tropical and temperate areas with an aridity index of less than 0.65. Drylands can be classified into four sub-types:
- Dry sub-humid lands
- Semi-arid lands
- Arid lands
- Hyper-arid lands
Some authorities regard hyper-arid lands as deserts (United Nations Convention to Combat Desertification) although a number of the world's deserts include both hyper-arid and arid climate zones. The UNCCD excludes hyper-arid zones from its definition of drylands.