Microbial mat in the context of "Prokaryote"

Marine life, sea life or ocean life is the collective ecological communities that encompass all aquatic animals, plants, algae, fungi, protists, single-celled microorganisms and associated viruses living in the saline water of marine habitats, either the sea water of marginal seas and oceans, or the brackish water of coastal wetlands, lagoons, estuaries and inland seas. As of 2023, more than 242,000 marine species have been documented, and perhaps two million marine species are yet to be documented. An average of 2,332 new species per year are being described. Marine life is studied scientifically in both marine biology and in biological oceanography.

By volume, oceans provide about 90% of the living space on Earth, and served as the cradle of life and vital biotic sanctuaries throughout Earth's geological history. The earliest known life forms evolved as anaerobic prokaryotes (archaea and bacteria) in the Archean oceans around the deep sea hydrothermal vents, before photoautotrophs appeared and allowed the microbial mats to expand into shallow water marine environments. The Great Oxygenation Event of the early Proterozoic significantly altered the marine chemistry, which likely caused a widespread anaerobe extinction event but also led to the evolution of eukaryotes through symbiogenesis between surviving anaerobes and aerobes. Complex life eventually arose out of marine eukaryotes during the Neoproterozoic, and which culminated in a large evolutionary radiation event of mostly sessile macrofaunae known as the Avalon Explosion. This was followed in the early Phanerozoic by a more prominent radiation event known as the Cambrian Explosion, where actively moving eumetazoan became prevalent. These marine life also expanded into fresh waters, where fungi and green algae that were washed ashore onto riparian areas started to take hold later during the Ordovician before rapidly expanding inland during the Silurian and Devonian, paving the way for terrestrial ecosystems to develop.

Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. According to modern evolutionary biology, all living beings could be descendants of a unique ancestor commonly referred to as the last universal common ancestor (LUCA) of all life on Earth.

Common descent is an effect of speciation, in which multiple species derive from a single ancestral population. The more recent the ancestral population two species have in common, the more closely they are related. The most recent common ancestor of all currently living organisms is the last universal ancestor, which lived about 3.9 billion years ago. The two earliest pieces of evidence for life on Earth are graphite found to be biogenic in 3.7 billion-year-old metasedimentary rocks discovered in western Greenland and microbial mat fossils found in 3.48 billion-year-old sandstone discovered in Western Australia. All currently living organisms on Earth share a common genetic heritage, though the suggestion of substantial horizontal gene transfer during early evolution has led to questions about the monophyly (single ancestry) of life. 6,331 groups of genes common to all living animals have been identified; these may have arisen from a single common ancestor that lived 650 million years ago in the Precambrian.

Stromatolites (/stroʊˈmætəˌlaɪts, strə-/ stroh-MAT-ə-lytes, strə-) or stromatoliths (from Ancient Greek στρῶμα (strôma), GEN στρώματος (strṓmatos) 'layer, stratum' and λίθος (líthos) 'rock') are layered sedimentary formations (microbialite) that are created mainly by photosynthetic microorganisms such as cyanobacteria, sulfate-reducing bacteria, and Pseudomonadota (formerly proteobacteria). These microorganisms produce adhesive compounds that cement sand and other rocky materials to form mineral "microbial mats". In turn, these mats build up layer by layer, growing gradually over time.

This process generates the characteristic lamination of stromatolites, a feature that is hard to interpret, in terms of its temporal and environmental significance. Different styles of stromatolite lamination have been described, which can be studied through microscopic and mathematical methods. A stromatolite may grow to a meter or more. Fossilized stromatolites provide important records of some of the most ancient life. As of the Holocene, living forms are rare.

Algal mats are one of many types of microbial mat that forms on the surface of water or rocks. They are typically composed of blue-green cyanobacteria and sediments. Formation occurs when alternating layers of blue-green bacteria and sediments are deposited or grow in place, creating dark-laminated layers. Stromatolites are prime examples of algal mats. Algal mats played an important role in the Great Oxidation Event on Earth some 2.3 billion years ago. Algal mats can become a significant ecological problem, if the mats grow so expansive or thick as to disrupt the other underwater marine life by blocking the sunlight or producing toxic chemicals.

Dickinsonia is a genus of extinct organism that lived during the late Ediacaran period in what is now Australia, China, Russia, and Ukraine. It had a round, approximately bilaterally symmetric body with multiple segments running along it. It could range from a few millimeters to over a meter in length, and likely lived in shallow waters, feeding on the microbial mats that dominated the seascape at the time.

As a member of the Ediacaran biota, its relationships to other organisms has been heavily debated. It was initially proposed to be a jellyfish, and over the years has been claimed to be a land-dwelling lichen, placozoan, or even a giant protist. Currently, the most popular interpretation is that it was a seafloor dwelling animal, perhaps a primitive stem group bilaterian, although this is still contentious. Among other Ediacaran organisms, it shares a close resemblance to other segmented forms like Vendia, Yorgia and Spriggina and has been proposed to be a member of the phylum Proarticulata or alternatively the morphogroup Dickinsoniomorpha. It is disputed whether the segments of Dickinsonia are bilaterally symmetric across the midline, or are offset from each other via glide reflection, or possibly both.