Hoof in the context of "Even-toed ungulate"

Ungulates are members of the diverse clade Euungulata, which primarily consists of large mammals with hooves. Once part of the taxon "Ungulata" along with paenungulates and tubulidentates, as well as several extinct taxa, "Ungulata" has since been determined to be a polyphyletic grouping based on molecular data. As a result, true ungulates had since been reclassified to the newer clade Euungulata in 2001 within the clade Laurasiatheria, while Paenungulata and Tubulidentata had been reclassified to the distant clade Afrotheria. Alternatively, some authors use the name Ungulata to designate the same clade as Euungulata.

Living ungulates are divided into two orders: Perissodactyla including equines, rhinoceroses, and tapirs; and Artiodactyla including cattle, antelope, pigs, giraffes, camels, sheep, deer, and hippopotamuses, among others. Cetaceans such as whales, dolphins, and porpoises are also classified as artiodactyls, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. Two other orders of ungulates, Notoungulata and Litopterna, both native to South America, became extinct at the end of the Pleistocene, around 12,000 years ago.

Footprints are the impressions or images left behind by a person walking or running. Hoofprints and pawprints are those left by animals with hooves or paws rather than feet, while "shoeprints" is the specific term for prints made by shoes. They may either be indentations in the ground or something placed onto the surface that was stuck to the bottom of the foot. A "trackway" is a set of footprints in soft earth left by a life-form; animal tracks are the footprints, hoofprints, or pawprints of an animal.

Footprints can be followed when tracking during a hunt or can provide evidence of activities. Some footprints remain unexplained, with several famous stories from mythology and legend. Others have provided evidence of prehistoric life and behaviours.

The evolution of the horse, a mammal of the family Equidae, occurred over a geologic time scale of 50 million years, transforming the small, dog-sized, forest-dwelling Eohippus into the large, single-toed, modern-day horse. Paleozoologists have been able to piece together a more complete outline of the evolutionary lineage of the modern horse than of any other animal. Much of this evolution took place in North America, where horses originated but became extinct about 10,000 years ago, before being reintroduced in the 15th century.

The horse belongs to the order Perissodactyla (odd-toed ungulates), the members of which share hooved feet and an odd number of toes on each foot, as well as mobile upper lips and a similar tooth structure. This means that horses share a common ancestry with tapirs and rhinoceroses. The perissodactyls arose in the late Paleocene, less than 10 million years after the Cretaceous–Paleogene extinction event. This group of animals appears to have been originally specialized for life in tropical forests, but whereas tapirs and, to some extent, rhinoceroses, retained their jungle specializations, modern horses are adapted to life in the climatic conditions of the steppes, which are drier and much harsher than forests or jungles. Other species of Equus are adapted to a variety of intermediate conditions.

In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals. The other options are digitigrade, walking on the toes and fingers with the heel and wrist permanently raised, and unguligrade, walking on the nail or nails of the toes (the hoof) with the heel/wrist and the digits permanently raised. The leg of a plantigrade mammal includes the bones of the upper leg (femur/humerus) and lower leg (tibia and fibula/radius and ulna). The leg of a digitigrade mammal also includes the metatarsals/metacarpals, the bones that in a human compose the arch of the foot and the palm of the hand. The leg of an unguligrade mammal also includes the phalanges, the finger and toe bones.

Among extinct animals, most early mammals such as pantodonts were plantigrade. A plantigrade foot is the primitive condition for mammals; digitigrade and unguligrade locomotion evolved later. Among archosaurs, the pterosaurs were partially plantigrade and walked on the whole of the hind foot and the fingers of the hand-wing. Out of the plantigrade animals, only a few, such as humans, kangaroos and certain rodents, are obligate bipeds, while most others are functional bipeds.

Keratin (/ˈkɛrətɪn/) is one of a family of structural fibrous proteins also known as scleroproteins. It is the key structural material making up scales, hair, nails, feathers, horns, claws, hooves, and the outer layer of skin in tetrapod vertebrates. Keratin also protects epithelial cells from damage or stress. Keratin is extremely insoluble in water and organic solvents. Keratin monomers assemble into bundles to form intermediate filaments, which are tough and form strong unmineralized epidermal appendages found in reptiles, birds, amphibians, and mammals. Excessive keratinization participate in fortification of certain tissues such as in horns of cattle and rhinos, and armadillos' osteoderm. The only other biological matter known to approximate the toughness of keratinized tissue is chitin.Keratin comes in two types: the primitive, softer forms found in all vertebrates and the harder, derived forms found only among sauropsids (reptiles and birds).

Artiodactyls are placental mammals belonging to the order Artiodactyla (/ˌɑːrtioʊˈdæktɪlə/ AR-tee-oh-DAK-tih-lə; from Ancient Greek ἄρτιος ártios 'even' and δάκτυλος dáktylos 'finger, toe'). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in the form of a hoof). The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, most perissodactyls bear weight on an odd number of the five toes. Another difference between the two orders is that many artiodactyls (except for Suina) digest plant cellulose in one or more stomach chambers rather than in their intestine (as perissodactyls do). Molecular biology, along with new fossil discoveries, has found that cetaceans (whales, dolphins, and porpoises) fall within this taxonomic branch, being most closely related to hippopotamuses. Some modern taxonomists thus apply the name Cetartiodactyla (/sɪˌtɑːrtioʊˈdæktɪlə/) to this group, while others opt to include cetaceans within the existing name of Artiodactyla. Some researchers use "even-toed ungulates" to exclude cetaceans and only include terrestrial artiodactyls, making the term paraphyletic in nature, but since cetaceans have 0 toes, and 0 is an even number, they technically fit the label.

The roughly 270 land-based even-toed ungulate species include pigs, peccaries, hippopotamuses, antelopes, deer, giraffes, camels, llamas, alpacas, sheep, goats and cattle. Many are herbivores, but suids are omnivorous, and cetaceans are entirely carnivorous. Artiodactyls are also known by many extinct groups such as anoplotheres, cainotheriids, merycoidodonts, entelodonts, anthracotheres, basilosaurids, and palaeomerycids. Many artiodactyls are of great dietary, economic, and cultural importance to humans.

Toes are the digits of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being digitigrade. Humans, and other animals that walk on the soles of their feet, are described as being plantigrade; unguligrade animals are those that walk on hooves at the tips of their toes.