Equidae in the context of "Ungulates"

The first evidence of horses in warfare dates from Eurasia between 4000 and 3000 BC. A Sumerian illustration of warfare from 2500 BC depicts some type of equine pulling wagons. By 1600 BC, improved harness and chariot designs made chariot warfare common throughout the Ancient Near East, and the earliest written training manual for war horses was a guide for training chariot horses written about 1350 BC. As formal cavalry tactics replaced the chariot, so did new training methods, and by 360 BC, the Greek cavalry officer Xenophon had written an extensive treatise on horsemanship. The effectiveness of horses in battle was also revolutionized by improvements in technology, such as the invention of the saddle, the stirrup, and the horse collar.

Many different types and sizes of horses were used in war, depending on the form of warfare. The type used varied with whether the horse was being ridden or driven, and whether they were being used for reconnaissance, cavalry charges, raiding, communication, or supply. Throughout history, mules and donkeys, as well as horses played a crucial role in providing support to armies in the field.

The horse (Equus ferus caballus) is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of Equus ferus. The horse has evolved over the past 45 to 55 million years from a small multi-toed creature, Eohippus, into the large single-toed animal of today, originally in North America. Humans began domesticating horses around 4000 BCE in Central Asia, and their domestication is believed to have been widespread by 3000 BCE. Horses in the subspecies caballus are domesticated, although some domesticated populations live in the wild as feral horses. These feral populations are not true wild horses, which are horses that have never been domesticated. There is an extensive, specialized vocabulary used to describe equine-related concepts, covering everything from anatomy to life stages, size, colors, markings, breeds, locomotion, and behavior.

Horses are adapted to run, allowing them to quickly escape predators, and possess a good sense of balance and a strong fight-or-flight response. Related to this need to flee from predators in the wild is an unusual trait: horses are able to sleep both standing up and lying down, with younger horses tending to sleep significantly more than adults. Female horses, called mares, carry their young for approximately 11 months and a young horse, called a foal, can stand and run shortly following birth. Most domesticated horses begin training under a saddle or in a harness between the ages of two and four. They reach full adult development by age five, and have an average lifespan of between 25 and 30 years.

A chariot is a type of vehicle similar to a cart, driven by a charioteer, usually using horses to provide rapid motive power. The oldest known chariots have been found in burials of the Sintashta culture in modern-day Chelyabinsk Oblast, Russia, dated to c. 1950–1880 BC and are depicted on cylinder seals from Central Anatolia in Kültepe dated to c. 1900 BC. The critical invention that allowed the construction of light, horse-drawn chariots was the spoked wheel.

The chariot was a fast, light, open, two-wheeled conveyance drawn by two or more equids (usually horses) that were hitched side by side, and was little more than a floor with a waist-high guard at the front and sides. It was initially used for ancient warfare during the Bronze and Iron Ages, but after its military capabilities had been superseded by light and heavy cavalries, chariots continued to be used for travel and transport, in processions, for games, and in races.

Equinae is a subfamily of the family Equidae, known from the Hemingfordian stage of the Early Miocene (16 million years ago) onwards. They originated in North America, before dispersing to every continent except Australia and Antarctica. They are thought to be a monophyletic grouping. Members of the subfamily are referred to as equines; the only extant equines are the horses, asses, and zebras of the genus Equus, with two other genera Haringtonhippus and Hippidion becoming extinct at the beginning of the Holocene, around 11–12,000 years ago.

The subfamily contains two tribes, the Equini and the Hipparionini, as well as two unplaced genera, Merychippus and Scaphohippus. Members of the family ancestrally had three toes, while members of the tribe Equini from the Middle Miocene onwards developed monodactyl feet.

Palaeotherium is an extinct genus of equoid that lived in Europe and possibly the Middle East from the Middle Eocene to the Early Oligocene. It is the type genus of the Palaeotheriidae, a group exclusive to the Palaeogene that was closest in relation to the Equidae, which contains horses plus their closest relatives and ancestors. Fossils of Palaeotherium were first described in 1782 by the French naturalist Robert de Lamanon and then closely studied by another French naturalist, Georges Cuvier, after 1798. Cuvier erected the genus in 1804 and recognized multiple species based on overall fossil sizes and forms. As one of the first fossil genera to be recognized with official taxonomic authority, it is recognized as an important milestone within the field of palaeontology. The research by early naturalists on Palaeotherium contributed to the developing ideas of evolution, extinction, and succession and demonstrating the morphological diversity of different species within one genus.

Since Cuvier's descriptions, many other naturalists from Europe and the Americas recognized many species of Palaeotherium, some valid, some reclassified to different genera afterward, and others being eventually rendered invalid. The German palaeontologist Jens Lorenz Franzen modernized its taxonomy due to his recognition of many subspecies as part of his dissertation in 1968, which were subsequently accepted by other palaeontologists. Today, there are sixteen known species recognized, many of which have multiple subspecies. In 1992, the French palaeontologist Jean-Albert Remy recognized two subgenera that most species are classified to based on cranial anatomies: the specialized Palaeotherium and the more generalized Franzenitherium.

Perissodactyla (/pəˌrɪsoʊˈdæktɪlə/, from Ancient Greek περισσός, perissós 'odd' and δάκτυλος, dáktylos 'finger, toe'), or odd-toed ungulates, is an order of ungulates. The order includes about 17 living species divided into three families: Equidae (horses, asses, and zebras), Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). They typically have reduced the weight-bearing toes to three or one of the five original toes, though tapirs retain four toes on their front feet. The nonweight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, artiodactyls (even-toed ungulates) bear most of their weight equally on four or two (an even number) of the five toes: their third and fourth toes. Another difference between the two is that perissodactyls digest plant cellulose in their intestines, rather than in one or more stomach chambers as artiodactyls, with the exception of Suina, do.

The order was considerably more diverse in the past, with notable extinct groups including the brontotheres, palaeotheres, chalicotheres, and the paraceratheres, with the paraceratheres including the largest known land mammals to have ever existed.