The thorax is the midsection (tagma) of the hexapod body (insects and entognathans). It holds the head, legs, wings and abdomen. It is also called mesosoma or cephalothorax in other arthropods.
It is formed by the prothorax, mesothorax and metathorax and comprises the scutellum; the cervix, a membrane that separates the head from the thorax; and the pleuron, a lateral sclerite of the thorax.
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In biology, a tagma (Greek: τάγμα, pl.: tagmata – τάγματα - body of soldiers; battalion) is a specialized grouping of multiple segments or metameres into a coherently functional morphological unit. Familiar examples are the head, the thorax, and the abdomen of insects. The segments within a tagma may be either fused (such as in the head of an insect) or so jointed as to be independently moveable (such as in the abdomen of most insects).
Usually the term is taken to refer to tagmata in the morphology of members of the phylum Arthropoda, but it applies equally validly in other phyla, such as the Chordata.
The subphylum Hexapoda (from Greek for 'six legs') or hexapods comprises the largest clade of arthropods and includes most of the extant arthropod species. It includes the crown group class Insecta (true insects), as well as the much smaller clade Entognatha, which includes three classes of wingless arthropods that were once considered insects: Collembola (springtails), Protura (coneheads) and Diplura (two-pronged bristletails). The insects and springtails are very abundant and are some of the most important pollinators, basal consumers, scavengers/detritivores and micropredators in terrestrial environments.
Hexapods are named for their most distinctive feature: a three-part body plan with a consolidated thorax and three pairs of legs. Most other arthropods have more than three pairs of legs. Most recent studies have recovered Hexapoda as a subgroup of Pancrustacea.
The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms prosoma and opisthosoma are equivalent to cephalothorax and abdomen in some groups. The terms prosoma and opisthosoma may be preferred by some researchers in cases such as arachnids, where there is neither fossil nor embryonic evidence animals in this class have ever had separate heads and thoraxes, and where the opisthosoma contains organs atypical of a true abdomen, such as a heart and respiratory organs.) The word cephalothorax is derived from the Greek words for head (κεφαλή, kephalé) and thorax (θώραξ, thórax). This fusion of the head and thorax is seen in chelicerates and crustaceans; in other groups, such as the Hexapoda (including insects), the head remains free of the thorax. In horseshoe crabs and many crustaceans, a hard shell called the carapace covers the cephalothorax.
Brachyura (from Ancient Greek βραχύς (brakhús), meaning "short", and οὐρά (ourá), meaning "tail"), is an infraorder of decapod crustaceans comprising the true crabs. They typically have a very short projecting tail-like abdomen, usually hidden entirely under the thorax. Their exoskeleton is often thickened and hard. They generally have five pairs of legs, and they have "pincers" or "claws" on the ends of the frontmost pair, scientifically termed the chelae. They are present in all the world's oceans, in freshwater, and on land, often hiding themselves in small crevices or burrowing into sediment. True crabs are omnivores, feeding on a variety of food, including a significant proportion of algae, as well as detritus and other invertebrates. True crabs are widely consumed by humans as food, with over 1.5 million tonnes caught annually.
True crabs first appeared in the fossil record during the Jurassic period, around 200 million years ago, achieving great diversity by the Cretaceous period; around 7,000 extant species in 96 families are known. The true crabs are the largest of the crab decapod groups with carcinised body forms. Smaller groups include the porcelain crabs, king crabs, and hairy stone crabs which have independently evolved similar forms and lifestyles.
The mesothorax is the middle of the three segments of the thorax of hexapods, and bears the second pair of legs. Its principal sclerites (exoskeletal plates) are the mesonotum (dorsal), the mesosternum (ventral), and the mesopleuron (lateral) on each side. The mesothorax is the segment that bears the forewings in all winged insects, though sometimes these may be reduced or modified, as in beetles (Coleoptera) or Dermaptera, in which they are sclerotized to form the elytra ("wing covers"), and the Strepsiptera, in which they are reduced to form halteres that attach to the mesonotum. All adult insects possess legs on the mesothorax. In some groups of insects, the mesonotum is hypertrophied, such as in Diptera, Hymenoptera, and Lepidoptera), in which the anterior portion of the mesonotum (called the mesoscutum, or simply "scutum") forms most of the dorsal surface of the thorax. In these orders, there is also typically a small sclerite attached to the mesonotum that covers the wing base, called the tegula. In one group of insects, the Hemiptera, the dorsal surface of the thorax is typically formed primarily of the prothorax, but also in part by the enlarged posterior portion of the mesonotum, called the scutellum; in the Coleoptera, the scutellum may or may not be visible, usually as a small triangular plate between the elytral bases, thus similar in position to the Hemipteran scutellum. In Diptera and Hymenoptera the mesothoracic scutellum is also distinct, but much smaller than the mesoscutum.