The Selfish Gene is a 1976 book by ethologist Richard Dawkins that promotes the gene-centered view of evolution, as opposed to views focused on the organism and the group. The book builds upon the thesis of George C. Williams's Adaptation and Natural Selection (1966); it also popularized ideas developed during the 1960s by W. D. Hamilton and others. From the gene-centred view, it follows that the more two individuals are genetically related, the more sense (at the level of the genes) it makes for them to behave cooperatively with each other.
A lineage is expected to evolve to maximise its inclusive fitnessβthe number of copies of its genes passed on globally (rather than by a particular individual). As a result, populations will tend towards an evolutionarily stable strategy. The book also introduces the term meme for a unit of human cultural evolution analogous to the gene, suggesting that such "selfish" replication may also model human culture, in a different sense. Memetics has become the subject of many studies since the publication of the book. In raising awareness of Hamilton's ideas, as well as making its own valuable contributions to the field, the book has also stimulated research on human inclusive fitness.