Polypodiopsida in the context of Hornwort

The order Cyatheales, which includes most tree ferns, is a taxonomic order of the fern class, Polypodiopsida. No clear morphological features characterize all of the Cyatheales, but DNA sequence data indicate the order is monophyletic. Some species in the Cyatheales have tree-like growth forms from a vertical rhizome, others have shorter or horizontal expanding rhizomes.

Some species have scales on the stems and leaves, while others have hairs. However, most plants in the Cyatheales are tree ferns and have trunk-like stems up to 10 metres (33 ft) tall. It is unclear how many times the tree form has evolved and been lost in the order.

The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (land plants with vascular tissues such as xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from non-vascular plants (mosses, hornworts and liverworts) by having specialized transport bundles that conduct water and nutrients from and to the roots, as well as life cycles in which the branched sporophyte is the dominant phase.

Ferns have complex leaves called megaphylls that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns that produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns and ophioglossoid ferns.

The order Salviniales (formerly known as the Hydropteridales and including the former Marsileales) is an order of ferns in the class Polypodiopsida.