Photosynthesis in the context of Photosystem II

The ocean is the body of salt water that covers approximately 70.8% of Earth. The ocean is conventionally divided into large bodies of water, which are also referred to as oceans (in descending order: the Pacific Ocean, the Atlantic Ocean, the Indian Ocean, the Antarctic/Southern Ocean, and the Arctic Ocean), and are themselves mostly divided into seas, gulfs and subsequent bodies of water. The ocean contains 97% of Earth's water and is the primary component of Earth's hydrosphere, acting as a huge reservoir of heat for Earth's energy budget, as well as for its carbon cycle and water cycle, forming the basis for climate and weather patterns worldwide. The ocean is essential to life on Earth, harbouring most of Earth's animals and protist life, originating photosynthesis and therefore Earth's atmospheric oxygen, still supplying half of it.

Ocean scientists split the ocean into vertical and horizontal zones based on physical and biological conditions. Horizontally the ocean covers the oceanic crust, which it shapes. Where the ocean meets dry land it covers relatively shallow continental shelfs, which are part of Earth's continental crust. Human activity is mostly coastal with high negative impacts on marine life. Vertically the pelagic zone is the open ocean's water column from the surface to the ocean floor. The water column is further divided into zones based on depth and the amount of light present. The photic zone starts at the surface and is defined to be "the depth at which light intensity is only 1% of the surface value" (approximately 200 m in the open ocean). This is the zone where photosynthesis can occur. In this process plants and microscopic algae (free-floating phytoplankton) use light, water, carbon dioxide, and nutrients to produce organic matter. As a result, the photic zone is the most biodiverse and the source of the food supply which sustains most of the ocean ecosystem. Light can only penetrate a few hundred more meters; the rest of the deeper ocean is cold and dark (these zones are called mesopelagic and aphotic zones).

A botanical name is a formal scientific name conforming to the International Code of Nomenclature for algae, fungi, and plants (ICN) and, if it concerns a plant cultigen, the additional cultivar or Group epithets must conform to the International Code of Nomenclature for Cultivated Plants (ICNCP). The code of nomenclature covers "all organisms traditionally treated as algae, fungi, or plants, whether fossil or non-fossil, including blue-green algae (Cyanobacteria), chytrids, oomycetes, slime moulds and photosynthetic protists with their taxonomically related non-photosynthetic groups (but excluding Microsporidia)."

The purpose of a formal name is to have a single name that is accepted and used worldwide for a particular plant or plant group. For example, the botanical name Bellis perennis denotes a plant species which is native to most of the countries of Europe and the Middle East, where it has accumulated various names in many languages. Later, the plant was introduced worldwide, bringing it into contact with more languages. English names for this plant species include: daisy, English daisy, and lawn daisy. The cultivar Bellis perennis 'Aucubifolia' is a golden-variegated horticultural selection of this species.

A leaf (pl.: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, flower, and fruit collectively form the shoot system. In most leaves, the primary photosynthetic tissue is the palisade mesophyll and is located on the upper side of the blade or lamina of the leaf, but in some species, including the mature foliage of Eucalyptus, palisade mesophyll is present on both sides and the leaves are said to be isobilateral. The leaf is an integral part of the stem system, and most leaves are flattened and have distinct upper (adaxial) and lower (abaxial) surfaces that differ in color, hairiness, the number of stomata (pores that intake and output gases), the amount and structure of epicuticular wax, and other features. Leaves are mostly green in color due to the presence of a compound called chlorophyll which is essential for photosynthesis as it absorbs light energy from the Sun. A leaf with lighter-colored or white patches or edges is called a variegated leaf.

Leaves vary in shape, size, texture and color, depending on the species The broad, flat leaves with complex venation of flowering plants are known as megaphylls and the species that bear them (the majority) as broad-leaved or megaphyllous plants, which also include acrogymnosperms and ferns. In the lycopods, with different evolutionary origins, the leaves are simple (with only a single vein) and are known as microphylls. Some leaves, such as bulb scales, are not above ground. In many aquatic species, the leaves are submerged in water. Succulent plants often have thick juicy leaves, but some leaves are without major photosynthetic function and may be dead at maturity, as in some cataphylls and spines. Furthermore, several kinds of leaf-like structures found in vascular plants are not totally homologous with them. Examples include flattened plant stems called phylloclades and cladodes, and flattened leaf stems called phyllodes which differ from leaves both in their structure and origin. Some structures of non-vascular plants look and function much like leaves. Examples include the phyllids of mosses and liverworts.

Marine microorganisms are defined by their habitat as microorganisms living in a marine environment, that is, in the saltwater of a sea or ocean or the brackish water of a coastal estuary. A microorganism (or microbe) is any microscopic living organism or virus, which is invisibly small to the unaided human eye without magnification. Microorganisms are very diverse. They can be single-celled or multicellular and include bacteria, archaea, viruses, and most protozoa, as well as some fungi, algae, and animals, such as rotifers and copepods. Many macroscopic animals and plants have microscopic juvenile stages. Some microbiologists also classify viruses as microorganisms, but others consider these as non-living.

Marine microorganisms have been variously estimated to make up between 70 and 90 percent of the biomass in the ocean. Taken together they form the marine microbiome. Over billions of years this microbiome has evolved many life styles and adaptations and come to participate in the global cycling of almost all chemical elements. Microorganisms are crucial to nutrient recycling in ecosystems as they act as decomposers. They are also responsible for nearly all photosynthesis that occurs in the ocean, as well as the cycling of carbon, nitrogen, phosphorus and other nutrients and trace elements. Marine microorganisms sequester large amounts of carbon and produce much of the world's oxygen.

Marine prokaryotes are marine bacteria and marine archaea. They are defined by their habitat as prokaryotes that live in marine environments, that is, in the saltwater of seas or oceans or the brackish water of coastal estuaries. All cellular life forms can be divided into prokaryotes and eukaryotes. Eukaryotes are organisms whose cells have a nucleus enclosed within membranes, whereas prokaryotes are the organisms that do not have a nucleus enclosed within a membrane. The three-domain system of classifying life adds another division: the prokaryotes are divided into two domains of life, the microscopic bacteria and the microscopic archaea, while everything else, the eukaryotes, become the third domain.

Prokaryotes play important roles in ecosystems as decomposers recycling nutrients. Some prokaryotes are pathogenic, causing disease and even death in plants and animals. Marine prokaryotes are responsible for significant levels of the photosynthesis that occurs in the ocean, as well as significant cycling of carbon and other nutrients.

An aquatic animal is any animal, whether vertebrate or invertebrate, that lives in a body of water for all or most of its lifetime. Aquatic animals generally conduct aquatic respiration by extracting dissolved oxygen in water via specialised respiratory organs called gills, through the skin or across enteral mucosae, although some are secondarily aquatic animals (e.g. marine reptiles and marine mammals) evolved from terrestrial ancestors that re-adapted to aquatic environments, in which case they actually use lungs to breathe air and are essentially holding their breath when living in water. Some species of gastropod mollusc, such as the eastern emerald sea slug, are even capable of kleptoplastic photosynthesis via endosymbiosis with ingested yellow-green algae.

Almost all aquatic animals reproduce in water, either oviparously or viviparously, and many species routinely migrate between different water bodies during their life cycle. Some animals have fully aquatic life stages (typically as eggs and larvae), while as adults they become terrestrial or semi-aquatic after undergoing metamorphosis. Such examples include amphibians such as frogs, many flying insects such as mosquitoes, mayflies, dragonflies, damselflies and caddisflies, as well as some species of cephalopod molluscs such as the algae octopus (whose larvae are completely planktonic, but adults are highly terrestrial).

Aquatic plants, also referred to as hydrophytes, are vascular plants and non-vascular plants that have adapted to live in aquatic environments (saltwater or freshwater). In lakes, rivers and wetlands, aquatic vegetations provide cover for aquatic animals such as fish, amphibians and aquatic insects, create substrate for benthic invertebrates, produce oxygen via photosynthesis, and serve as food for some herbivorous wildlife. Familiar examples of aquatic plants include waterlily, lotus, duckweeds, mosquito fern, floating heart, water milfoils, mare's tail, water lettuce, water hyacinth, and algae.

Aquatic plants require special adaptations for prolonged inundation in water, and for floating at the water surface. The most common adaptation is the presence of lightweight internal packing cells, aerenchyma, but floating leaves and finely dissected leaves are also common. Aquatic plants only thrive in water or in soil that is frequently saturated, and are therefore a common component of swamps and marshlands.

Algae (/ˈældʒiː/ AL-jee, UK also /ˈælɡiː/ AL-ghee; sg.: alga /ˈælɡə/ AL-gə) is an informal term for any organisms of a large and diverse group of photosynthetic organisms that are not land plants, and includes species from multiple distinct clades. Such organisms range from unicellular microalgae, such as cyanobacteria, Chlorella, and diatoms, to multicellular macroalgae such as kelp or brown algae which may grow up to 50 metres (160 ft) in length. Most algae are aquatic organisms and lack many of the distinct cell and tissue types, such as stomata, xylem, and phloem that are found in land plants. The largest and most complex marine algae are called seaweeds. In contrast, the most complex freshwater forms are the Charophyta, a division of green algae which includes, for example, Spirogyra and stoneworts. Algae that are carried passively by water are plankton, specifically phytoplankton.

Algae constitute a polyphyletic group because they do not include a common ancestor, and although eukaryotic algae with chlorophyll-bearing plastids seem to have a single origin (from symbiogenesis with cyanobacteria), they were acquired in different ways. Green algae are a prominent example of algae that have primary chloroplasts derived from endosymbiont cyanobacteria. Diatoms and brown algae are examples of algae with secondary chloroplasts derived from endosymbiotic red algae, which they acquired via phagocytosis. Algae exhibit a wide range of reproductive strategies, from simple asexual cell division to complex forms of sexual reproduction via spores.

Photoautotrophs are organisms that can utilize light energy from sunlight, and elements (such as carbon) from inorganic compounds, to produce organic materials needed to sustain their own metabolism (i.e. autotrophy). Such biological activities are known as photosynthesis, and examples of such organisms include plants, algae and cyanobacteria.

Eukaryotic photoautotrophs absorb photonic energy through the photopigment chlorophyll (a porphyrin derivative) in their endosymbiont chloroplasts, while prokaryotic photoautotrophs use chlorophylls and bacteriochlorophylls present in free-floating cytoplasmic thylakoids. Plants, algae, and cyanobacteria perform oxygenic photosynthesis that produces oxygen as a byproduct, while some bacteria perform anoxygenic photosynthesis.

The Great Oxidation Event (GOE) or Great Oxygenation Event, also called the Oxygen Catastrophe, Oxygen Revolution, Oxygen Crisis, or Oxygen Holocaust, was a time interval during the Earth's Paleoproterozoic era when the Earth's atmosphere and shallow seas first experienced a rise in the concentration of free oxygen. This began approximately 2.460–2.426 billion years ago (Ga) during the Siderian period and ended approximately 2.060 Ga ago during the Rhyacian. Geological, isotopic and chemical evidence suggests that biologically produced molecular oxygen (dioxygen or O₂) started to accumulate in the Archean prebiotic atmosphere due to microbial photosynthesis, and eventually changed it from a weakly reducing atmosphere practically devoid of oxygen into an oxidizing one containing abundant free oxygen, with oxygen levels being as high as 10% of modern atmospheric level by the end of the GOE.

The green algae (sg.: green alga) are a group of chlorophyll-containing autotrophic algae consisting of the phylum Prasinodermophyta and its unnamed sister group that contains the Chlorophyta and Charophyta/Streptophyta. The land plants (Embryophyta) have emerged deep within the charophytes as a sister of the Zygnematophyceae. Since the realization that the Embryophyta emerged within the green algae, some authors are starting to include them. The completed clade that includes both green algae and embryophytes is monophyletic and is referred to as the clade Viridiplantae and as the kingdom Plantae. The green algae include unicellular and colonial flagellates, most with two flagella per cell, as well as various colonial, coccoid (spherical), and filamentous forms, and macroscopic, multicellular seaweeds. There are about 22,000 species of green algae, many of which live most of their lives as single cells, while other species form coenobia (colonies), long filaments, or highly differentiated macroscopic seaweeds.

A few other organisms rely on green algae to conduct photosynthesis for them. The chloroplasts in dinoflagellates of the genus Lepidodinium, euglenids and chlorarachniophytes were acquired from ingested endosymbiont green algae, and in the latter retain a nucleomorph (vestigial nucleus). Green algae are also found symbiotically in the ciliate Paramecium, and in Hydra viridissima and in flatworms. Some species of green algae, particularly of genera Trebouxia of the class Trebouxiophyceae and Trentepohlia (class Ulvophyceae), can be found in symbiotic associations with fungi to form lichens. In general, the fungal species that partner in lichens cannot live on their own, while the algal species is often found living in nature without the fungus. Trentepohlia is a filamentous green alga that can live independently on humid soil, rocks or tree bark or form the photosymbiont in lichens of the family Graphidaceae. Also the macroalga Prasiola calophylla (Trebouxiophyceae) is terrestrial, andPrasiola crispa, which live in the supralittoral zone, is terrestrial and can in the Antarctic form large carpets on humid soil, especially near bird colonies.

Plants are the eukaryotes that comprise the kingdom Plantae; they are predominantly photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with cyanobacteria to produce sugars from carbon dioxide and water, using the green pigment chlorophyll. Exceptions are parasitic plants that have lost the genes for chlorophyll and photosynthesis, and obtain their energy from other plants or fungi. Most plants are multicellular, except for some green algae.

Historically, as in Aristotle's biology, the plant kingdom encompassed all living things that were not animals, and included algae and fungi. Definitions have narrowed since then; current definitions exclude fungi and some of the algae. By the definition used in this article, plants form the clade Viridiplantae (green plants), which consists of the green algae and the embryophytes or land plants (hornworts, liverworts, mosses, lycophytes, ferns, conifers and other gymnosperms, and flowering plants). A definition based on genomes includes the Viridiplantae, along with the red algae and the glaucophytes, in the clade Archaeplastida.

Natural gas (also methane gas, and gas) is a naturally occurring compound of gaseous hydrocarbons, primarily methane (95%), small amounts of higher alkanes, and traces of carbon dioxide and nitrogen, hydrogen sulfide and helium. Methane is a colorless and odorless gas, and, after carbon dioxide, is the second-greatest greenhouse gas that contributes to global climate change. Because natural gas is odorless, a commercial odorizer, such as methanethiol, that smells of hydrogen sulfide (rotten eggs) is added to the gas for the ready detection of gas leaks.

Natural gas is a fossil fuel that is formed when layers of organic matter (primarily marine microorganisms) are thermally decomposed under oxygen-free conditions, subjected to intense heat and pressure underground over millions of years. The energy that the decayed organisms originally obtained from the sun via photosynthesis is stored as chemical energy within the molecules of methane and other hydrocarbons.

An algal bloom or algae bloom is a rapid increase or accumulation in the population of algae in fresh water or marine water systems. It may be a benign or harmful algal bloom.

Algal bloom is often recognized by the discoloration in the water from the algae's pigments. The term algae encompasses many types of aquatic photosynthetic organisms, both macroscopic multicellular organisms like seaweed and microscopic unicellular organisms like cyanobacteria. Algal bloom commonly refers to the rapid growth of microscopic unicellular algae, not macroscopic algae. An example of a macroscopic algal bloom is a kelp forest.

The photic zone (or euphotic zone, epipelagic zone, or sunlight zone) is the uppermost layer of a body of water that receives sunlight, allowing phytoplankton to perform photosynthesis. It undergoes a series of physical, chemical, and biological processes that supply nutrients into the upper water column. The photic zone is home to the majority of aquatic life due to the activity (primary production) of the phytoplankton. The thicknesses of the photic and euphotic zones vary with the intensity of sunlight as a function of season and latitude and with the degree of water turbidity. The bottommost, or aphotic, zone is the region of perpetual darkness that lies beneath the photic zone and includes most of the ocean waters.