Palynology in the context of "Climate proxy"

Marker horizons (also referred to as chronohorizons, key beds or marker beds) are stratigraphic units of the same age and of such distinctive composition and appearance, that, despite their presence in separate geographic locations, there is no doubt about their being of equivalent age (isochronous) and of common origin. Such clear markers facilitate the correlation of strata, and used in conjunction with fossil floral and faunal assemblages and paleomagnetism, permit the mapping of land masses and bodies of water throughout the history of the earth. They usually consist of a relatively thin layer of sedimentary rock that is readily recognized on the basis of either its distinct physical characteristics or fossil content and can be mapped over a very large geographic area. As a result, a key bed is useful for correlating sequences of sedimentary rocks over a large area. Typically, key beds were created as the result of either instantaneous events or (geologically speaking) very short episodes of the widespread deposition of a specific types of sediment. As the result, key beds often can be used for both mapping and correlating sedimentary rocks and dating them. Volcanic ash beds (tonsteins and bentonite beds) and impact spherule beds, and specific megaturbidites are types of key beds created by instantaneous events. The widespread accumulation of distinctive sediments over a geologically short period of time have created key beds in the form of peat beds, coal beds, shell beds, marine bands, black shales in cyclothems, and oil shales. A well-known example of a key bed is the global layer of iridium-rich impact ejecta that marks the Cretaceous–Paleogene boundary (K–T boundary).

Palynology, the study of fossil pollens and spores, routinely works out the stratigraphy of rocks by comparing pollen and spore assemblages with those of well-known layers—a tool frequently used by petroleum exploration companies in the search for new fields. The fossilised teeth or elements of conodonts are an equally useful tool.

Transmission electron microscopy (TEM) is a microscopy technique in which a beam of electrons is transmitted through a specimen to form an image. The specimen is most often an ultrathin section less than 100 nm thick or a suspension on a grid. An image is formed from the interaction of the electrons with the sample as the beam is transmitted through the specimen. The image is then magnified and focused onto an imaging device, such as a fluorescent screen, a layer of photographic film, or a detector such as a scintillator attached to a charge-coupled device or a direct electron detector.

Transmission electron microscopes are capable of imaging at a significantly higher resolution than light microscopes, owing to the smaller de Broglie wavelength of electrons. This enables the instrument to capture fine detail—even as small as a single column of atoms, which is thousands of times smaller than a resolvable object seen in a light microscope. Transmission electron microscopy is a major analytical method in the physical, chemical and biological sciences. TEMs find application in cancer research, virology, and materials science as well as pollution, nanotechnology and semiconductor research, but also in other fields such as paleontology and palynology.

Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells).

Pollen grains have a hard coat made of sporopollenin that protects the gametophytes during the process of their movement from the stamens to the pistil of flowering plants, or from the male cone to the female cone of gymnosperms. If pollen lands on a compatible pistil or female cone, it germinates, producing a pollen tube that transfers the sperm to the ovule containing the female gametophyte. Individual pollen grains are small enough to require magnification to see detail. The study of pollen is called palynology and is highly useful in paleoecology, paleontology, archaeology, and forensics.

Paleobotany or palaeobotany, also known as paleophytology, is the branch of botany dealing with the recovery and identification of plant fossils from geological contexts, and their use for the biological reconstruction of past environments (paleogeography), and the evolutionary history of plants, with a bearing upon the evolution of life in general. It is a component of paleontology and paleobiology. The prefix palaeo- or paleo- means "ancient, old", and is derived from the Greek adjective παλαιός, palaios. Paleobotany includes the study of land plants, as well as the study of prehistoric marine photoautotrophs such as photosynthetic algae, seaweeds or kelp. A closely related field is palynology, which is the study of fossilized and extant spores and pollen.

Paleobotany is important in the reconstruction of ancient ecological and climate systems, known as paleoecology and paleoclimatology respectively. It is fundamental to the study of green plant development and evolution. Paleobotany is a historical science much like its adjacent, paleontology. Because of the understanding that paleobotany gives to archeologists, it has become important to the field of archaeology as a whole. primarily for the use of phytoliths in relative dating and in paleoethnobotany.