Keystone species in the context of "Saguaro"

A beaver dam or beaver impoundment is a dam built by beavers; it creates a pond which protects against predators and holds food during winter. These structures modify the natural environment in such a way that the overall ecosystem builds upon the change, making beavers a keystone species and ecosystem engineers. They build prolifically at night, carrying mud with their forepaws and timber between their teeth.

Beavers (genus Castor) are large, semiaquatic rodents of the Northern Hemisphere. There are two existing species: the North American beaver (Castor canadensis) and the Eurasian beaver (C. fiber). Beavers are the second-largest living rodents, after capybaras, weighing up to 50 kg (110 lb). They have stout bodies with large heads, long chisel-like incisors, brown or gray fur, hand-like front feet, webbed back feet, and tails that are flat and scaly. The two species differ in skull and tail shape and fur color. Beavers can be found in a number of freshwater habitats, such as rivers, streams, lakes and ponds. They are herbivorous, consuming tree bark, aquatic plants, grasses and sedges.

Beavers build dams and lodges using tree branches, vegetation, rocks and mud; they chew down trees for building material. Dams restrict water flow, forming ponds, and lodges (usually built in ponds) serve as shelters. Their infrastructure creates wetlands used by many other species, and because of their effect on other organisms in the ecosystem, beavers are considered a keystone species. Adult males and females live in monogamous pairs with their offspring. After their first year, the young help their parents repair dams and lodges; older siblings may also help raise newly born offspring. Beavers hold territories and mark them using scent mounds made of mud, debris, and castoreum—a liquid substance excreted through the beaver's urethra-based castor sacs. Beavers can also recognize their kin by their anal gland secretions and are more likely to tolerate them as neighbors.

An ecosystem engineer is any species that creates, significantly modifies, maintains or destroys a habitat. These organisms can have a large impact on species richness and landscape-level heterogeneity of an area. As a result, ecosystem engineers are important for maintaining the health and stability of the environment they are living in. Since all organisms impact the environment they live in one way or another, it has been proposed that the term "ecosystem engineers" be used only for keystone species whose behavior very strongly affects other organisms.

The walrus (Odobenus rosmarus) is a large pinniped marine mammal with discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. It is the only extant species in the family Odobenidae and genus Odobenus. This species is subdivided into two subspecies: the Atlantic walrus (O. r. rosmarus), which lives in the Atlantic Ocean, and the Pacific walrus (O. r. divergens), which lives in the Pacific Ocean.

Adult walrus are characterised by prominent tusks and whiskers, and considerable bulk: adult males in the Pacific can weigh more than 2,000 kilograms (4,400 pounds) and, among pinnipeds, are exceeded in size only by the two species of elephant seals. Walrus live mostly in shallow waters above the continental shelves, spending significant amounts of their lives on the sea ice looking for benthic bivalve molluscs. Walruses are relatively long-lived, social animals, and are considered to be a "keystone species" in the Arctic marine regions.

Destructive fishing practices are fishing practices which easily result in irreversible damage to habitats and the sustainability of the fishery ecosystems. Such damages can be caused by direct physical destruction of the underwater landform and vegetation, overfishing (especially of keystone species), indiscriminate killing/maiming of aquatic life, disruption of vital reproductive cycles, and lingering water pollution.

Many fishing techniques can be destructive if used inappropriately, but some practices (such as blasting, electrocution and poisoning) are particularly likely to result in irreversible damage to the ecosystem. These practices are mostly, though not always, illegal (see also illegal, unreported, and unregulated fishing), and even where they are illegal, regulations are often inadequately enforced.

An oak is a hardwood tree or shrub in the genus Quercus of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisphere, with some 500 species, both deciduous and evergreen. Fossil oaks date back to the Middle Eocene. Molecular phylogeny shows that the genus is divided into Old World and New World clades, but many oak species hybridise freely, making the genus's history difficult to resolve.

Ecologically, oaks are keystone species in habitats from Mediterranean semi-desert to subtropical rainforest. They live in association with many kinds of fungi including truffles. Oaks support more than 950 species of caterpillar, many kinds of gall wasp which form distinctive galls (roundish woody lumps such as the oak apple), and a large number of pests and diseases. Oak leaves and acorns contain enough tannin to be toxic to cattle, but pigs are able to digest them safely. Oak timber is strong and hard, and has found many uses in construction and furniture-making. The bark was traditionally used for tanning leather. Wine barrels are made of oak; these are used for aging alcoholic beverages such as sherry and whisky, giving them a range of flavours, colours, and aromas. The spongy bark of the cork oak is used to make traditional wine bottle corks. Almost a third of oak species are threatened with extinction due to climate change, invasive pests, and habitat loss.

In 1992, Northern cod populations fell to 1% of historic levels, in large part from decades of overfishing. The Canadian Federal Minister of Fisheries and Oceans, John Crosbie, declared a moratorium on the Northern Cod fishery, which had primarily shaped the lives and communities of Canada's eastern coast for 500 years. A significant factor contributing to the depletion of the cod stocks off Newfoundland's shores was the introduction of equipment and technology that increased landed fish volume. From the 1950s onwards, new technology allowed fishers to trawl a larger area more deeply and for longer, with the catches peaking in the 1970s and 1980s. Cod stocks were depleted at a faster rate than could be replenished.

The trawlers also caught enormous amounts of non-commercial fish, which were economically unimportant but very important ecologically. The incidental catch undermined the stability of the ecosystem by depleting stocks of important predator and prey species.

Megaherbivores (Greek μέγας megas "large" and Latin herbivora "herbivore") are large herbivores that can exceed 1,000 kg (2,200 lb) in weight. The earliest herbivores to reach such sizes like the pareiasaurs appeared in the Permian period. During most of the Mesozoic, the megaherbivore niche was largely dominated by dinosaurs up until their extinction during the Cretaceous–Paleogene extinction event. After this period, small mammalian species evolved into large herbivores in the Paleogene. As part of the Late Pleistocene megafauna extinctions, 80% of megaherbivore species became extinct, with megaherbivores becoming entirely extinct in Europe, Australia and the Americas. Recent megaherbivores include elephants, rhinos, hippos, and giraffes. There are nine extant species of terrestrial megaherbivores living in Africa and Asia. The African bush elephant is the largest extant species.

Extant megaherbivores are keystone species in their environment. They defoliate the landscape and spread a greater number of seeds than other frugivores. Extant megaherbivores, like most large mammals, are K-selected species and are characterized by their large size, relative immunity to predation, their effect on plant species, and their dietary tolerance.

Forage fish, also called prey fish or bait fish, are small pelagic fish that feed on planktons (i.e. planktivores) and other small aquatic organisms (e.g. krill). They are in turn preyed upon by various predators including larger fish, seabirds and marine mammals, this making them keystone species in their aquatic ecosystems.

The typical ocean forage fish feed at the lower trophic level of the food chain, often by filter feeding. They include particularly fishes of the order Clupeiformes (herrings, sardines, shad, hilsa, menhaden, anchovies, and sprats), but also other small fish, including halfbeaks, silversides, smelt such as capelin and goldband fusiliers.