Hornwort in the context of "Plant kingdom"

Plants are the eukaryotes that comprise the kingdom Plantae; they are predominantly photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with cyanobacteria to produce sugars from carbon dioxide and water, using the green pigment chlorophyll. Exceptions are parasitic plants that have lost the genes for chlorophyll and photosynthesis, and obtain their energy from other plants or fungi. Most plants are multicellular, except for some green algae.

Historically, as in Aristotle's biology, the plant kingdom encompassed all living things that were not animals, and included algae and fungi. Definitions have narrowed since then; current definitions exclude fungi and some of the algae. By the definition used in this article, plants form the clade Viridiplantae (green plants), which consists of the green algae and the embryophytes or land plants (hornworts, liverworts, mosses, lycophytes, ferns, conifers and other gymnosperms, and flowering plants). A definition based on genomes includes the Viridiplantae, along with the red algae and the glaucophytes, in the clade Archaeplastida.

Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (/braɪˈɒfətə/, /ˌbraɪ.əˈfaɪtə/) sensu stricto. Bryophyta (sensu lato, Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hornworts. Mosses typically form dense green clumps or mats, often in damp or shady locations. The individual plants are usually composed of simple leaves that are generally only one cell thick, attached to a stem that may be branched or unbranched and has only a limited role in conducting water and nutrients. Although some species have conducting tissues, these are generally poorly developed and structurally different from similar tissue found in vascular plants. Mosses do not have seeds and after fertilisation develop sporophytes with unbranched stalks topped with single capsules containing spores. They are typically 0.2–10 cm (0.1–3.9 in) tall, though some species are much larger. Dawsonia superba, the tallest moss in the world, can grow to 60 cm (24 in) in height. There are approximately 12,000 species.

Mosses are commonly confused with liverworts, hornworts and lichens. Although often described as non-vascular plants, many mosses have advanced vascular systems. Like liverworts and hornworts, the haploid gametophyte generation of mosses is the dominant phase of the life cycle. This contrasts with the pattern in all vascular plants (seed plants and pteridophytes), where the diploid sporophyte generation is dominant. Lichens may superficially resemble mosses, and sometimes have common names that include the word "moss" (e.g., "reindeer moss" or "Iceland moss"), but they are fungal symbioses and not related to mosses.

Gynoecium (/ɡaɪˈniːsi.əm, dʒɪˈniːʃi.əm/; from Ancient Greek γυνή (gunḗ) 'woman, female' and οἶκος (oîkos) 'house', pl. gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) pistils and is typically surrounded by the pollen-producing reproductive organs, the stamens, collectively called the androecium. The gynoecium is often referred to as the "female" portion of the flower, although rather than directly producing female gametes (i.e. egg cells), the gynoecium produces megaspores, each of which develops into a female gametophyte which then produces egg cells.

The term gynoecium is also used by botanists to refer to a cluster of archegonia and any associated modified leaves or stems present on a gametophyte shoot in mosses, liverworts, and hornworts. The corresponding terms for the male parts of those plants are clusters of antheridia within the androecium. Flowers that bear a gynoecium but no stamens are called pistillate or carpellate. Flowers lacking a gynoecium are called staminate.

The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (land plants with vascular tissues such as xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from non-vascular plants (mosses, hornworts and liverworts) by having specialized transport bundles that conduct water and nutrients from and to the roots, as well as life cycles in which the branched sporophyte is the dominant phase.

Ferns have complex leaves called megaphylls that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns that produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns and ophioglossoid ferns.

The embryophytes (/ˈɛmbriəˌfaɪts/) are a clade of plants, known as Embryophyta (Plantae sensu strictissimo) (/ˌɛmbriˈɒfətə, -oʊˈfaɪtə/) or land plants. They are the most familiar group of photoautotrophs that make up the vegetation on Earth's dry lands and wetlands. Embryophytes have a common ancestor with green algae, having emerged within the Phragmoplastophyta clade of freshwater charophyte green algae as a sister taxon of Charophyceae, Coleochaetophyceae and Zygnematophyceae. Embryophytes consist of the bryophytes and the polysporangiophytes. Living embryophytes include hornworts, liverworts, mosses, lycophytes, ferns, gymnosperms and angiosperms (flowering plants). Embryophytes have haplodiplontic life cycles.

The embryophytes are informally called "land plants" because they thrive primarily in terrestrial habitats (despite some members having evolved secondarily to live once again in semiaquatic/aquatic habitats), while the related green algae are primarily aquatic. Embryophytes are complex multicellular eukaryotes with specialized reproductive organs. The name derives from their innovative characteristic of nurturing the young embryo sporophyte during the early stages of its multicellular development within the tissues of the parent gametophyte. With very few exceptions, embryophytes obtain biological energy by photosynthesis, using chlorophyll a and b to harvest the light energy in sunlight for carbon fixation from carbon dioxide and water in order to synthesize carbohydrates while releasing oxygen as a byproduct. The study of land plants is called phytology.

Bryophytes (/ˈbraɪ.əˌfaɪts/) are a group of land plants (embryophytes), sometimes treated as a taxonomic division referred to as Bryophyta sensu lato, that contains three groups of non-vascular land plants: the liverworts, hornworts, and mosses. In the strict sense, the division Bryophyta consists of the mosses only. Bryophytes are characteristically limited in size and prefer moist habitats although some species can survive in drier environments. The bryophytes consist of about 20,000 plant species. Bryophytes produce enclosed reproductive structures (gametangia and sporangia), but they do not produce flowers or seeds. They reproduce sexually by spores and asexually by fragmentation or the production of gemmae.

Though bryophytes were considered a paraphyletic group in recent years, almost all of the most recent phylogenetic evidence supports the monophyly of this group, as originally classified by Wilhelm Schimper in 1879.

Plant reproductive morphology is the study of the physical form and structure (the morphology) of those parts of plants directly or indirectly concerned with sexual reproduction.

Among all living organisms, flowers, which are the reproductive structures of flowering plants (angiosperms), are the most varied physically and show a correspondingly great diversity in methods of reproduction. Plants that are not flowering plants (green algae, mosses, liverworts, hornworts, ferns and gymnosperms such as conifers) also have complex interplays between morphological adaptation and environmental factors in their sexual reproduction.

An archegonium (pl.: archegonia), from the Ancient Greek ἀρχή ("beginning") and γόνος ("offspring"), is a multicellular structure or organ of the gametophyte phase of certain plants, producing and containing the ovum or female gamete. The corresponding male organ is called the antheridium. The archegonium has a long neck canal or venter and a swollen base. Archegonia are typically located on the surface of the plant thallus, although in the hornworts they are embedded.