Halteres in the context of "Flies"

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⭐ Core Definition: Halteres

Halteres (/hælˈtɪərz/; singular halter or haltere) (from Ancient Greek: ἁλτῆρες, hand-held weights to give an impetus in leaping) are a pair of small club-shaped organs on the body of some flying insects that provide information about body rotations during flight. They are present on flies belonging to two orders, Diptera and Strepsiptera. In dipterans, the halteres evolved from a pair of ancestral hindwings, while males of the much smaller strepsiterans (stylops) have halteres evolved from a pair of ancestral forewings.

Halteres oscillate rapidly along with the wings and operate like vibrating structure gyroscopes: any rotation of the plane of oscillation causes a force on the vibrating halteres by the Coriolis effect. The insect detects this force with sensory organs called campaniform sensilla and chordotonal organs located at the base of the halteres and uses this information to interpret and correct its position in space. Halteres provide rapid feedback to the wing-steering muscles, as well as to the muscles responsible for stabilizing the head.

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👉 Halteres in the context of Flies

Flies are insects of the order Diptera, the name being derived from the Greek δι- di- "two", and πτερόν pteron "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing more than 150,000 species including horse-flies, crane flies, hoverflies, mosquitoes and others.

Flies have a mobile head, with a pair of large compound eyes, and mouthparts designed for piercing and sucking (mosquitoes, black flies and robber flies), or for lapping and sucking in the other groups. Their wing arrangement gives them great manoeuvrability in flight, and claws and pads on their feet enable them to cling to smooth surfaces. Flies undergo complete metamorphosis; the eggs are often laid on the larval food-source and the larvae, which lack true limbs, develop in a protected environment, often inside their food source. Other species are ovoviviparous, opportunistically depositing hatched or hatching larvae instead of eggs on carrion, dung, decaying material, or open wounds of mammals. The pupa is a tough capsule from which the adult emerges when ready to do so; flies mostly have short lives as adults.

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Halteres in the context of Morphology of Diptera

Dipteran morphology differs in some significant ways from the broader morphology of insects. The Diptera is a very large and diverse order of mostly small to medium-sized insects. They have prominent compound eyes on a mobile head, and (at most) one pair of functional, membraneous wings, which are attached to a complex mesothorax. The second pair of wings, on the metathorax, are reduced to halteres. The order's fundamental peculiarity is its remarkable specialization in terms of wing shape and the morpho-anatomical adaptation of the thorax – features which lend particular agility to its flying forms. The filiform, stylate or aristate antennae correlate with the Nematocera, Brachycera and Cyclorrhapha taxa respectively. It displays substantial morphological uniformity in lower taxa, especially at the level of genus or species. The configuration of integumental bristles is of fundamental importance in their taxonomy, as is wing venation. It displays a complete metamorphosis (egg, larva, pupa, adult), or holometabolous development. The larvae are legless, and have head capsules with mandibulate mouthparts in the Nematocera. The larvae of "higher flies" (Brachycera) are however headless and wormlike, and display only three instars. Pupae are obtect in the Nematocera, or coarcate in Brachycera.

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Halteres in the context of Vibrating structure gyroscope

A vibrating structure gyroscope (VSG), defined by the IEEE as a Coriolis vibratory gyroscope (CVG), is a gyroscope that uses a vibrating (as opposed to rotating) structure as its orientation reference. A vibrating structure gyroscope functions much like the halteres of flies (insects in the order Diptera).

The underlying physical principle is that a vibrating object tends to continue vibrating in the same plane even if its support rotates. The Coriolis effect causes the object to exert a force on its support, and by measuring this force the rate of rotation can be determined.

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