Fish fin in the context of "Flipper (anatomy)"

A fish is an aquatic, anamniotic, gill-bearing vertebrate animal with swimming fins and a hard skull, but lacking limbs with digits. Fish can be grouped into the more basal jawless fish and the more common jawed fish, the latter including all living cartilaginous and bony fish, as well as the extinct placoderms and acanthodians. In a break from the long tradition of grouping all fish into a single class (Pisces), modern phylogenetics views fish as a paraphyletic group which includes all vertebrates except tetrapods. In English, the plural of "fish" is fish when referring to individuals and fishes when referring to species.

Most fish are cold-blooded, their body temperature varying with the surrounding water, though some large, active swimmers like the white shark and tuna can maintain a higher core temperature. Many fish can communicate acoustically with each other, such as during courtship displays. The study of fish is known as ichthyology.

A limb (from Old English lim, meaning "body part") is a jointed, muscled appendage of a tetrapod vertebrate animal used for weight-bearing, terrestrial locomotion and physical interaction with other objects. The distalmost portion of a limb is known as its extremity. The limbs' bony endoskeleton, known as the appendicular skeleton, is homologous among all tetrapods, who use their limbs for walking, running and jumping, swimming, climbing, grasping, touching and striking.

All tetrapods have four limbs that are organized into two bilaterally symmetrical pairs, with one pair at each end of the torso, which phylogenetically correspond to the four paired fins (pectoral and pelvic fins) of their fish (sarcopterygian) ancestors. The cranial pair (i.e. closer to the head) of limbs are known as the forelimbs or front legs, and the caudal pair (i.e. closer to the tail or coccyx) are the hindlimbs or back legs. In animals with a more erect bipedal posture (mainly hominid primates, particularly humans), the forelimbs and hindlimbs are often called upper and lower limbs, respectively. The fore-/upper limbs are connected to the thoracic cage via the pectoral/shoulder girdles, and the hind-/lower limbs are connected to the pelvis via the hip joints. Many animals, especially the arboreal species, have prehensile forelimbs adapted for grasping and climbing, while some (mostly primates) can also use hindlimbs for grasping. Some animals (birds and bats) have expanded forelimbs (and sometimes hindlimbs as well) with specialized feathers or membranes to achieve lift and fly. Aquatic and semiaquatic tetrapods usually have limb features (such as webbings) adapted to better provide propulsion in water, while marine mammals and sea turtles have convergently evolved flattened, paddle-like limbs known as flippers.

Several groups of tetrapods have undergone secondary aquatic adaptation, an evolutionary transition from being purely terrestrial to living at least partly aquatic. These animals are called "secondarily aquatic" because although all tetrapods descended from freshwater lobe finned fish (see evolution of tetrapods), their more recent ancestors are terrestrial vertebrates that evolved on land for hundreds of millions of years, and their clades only re-adapted to aquatic environment much later.

Unlike primarily aquatic vertebrates (i.e. fish), secondarily aquatic tetrapods (especially aquatic amniotes), while having appendages such as flippers, dorsal fin and tail fins (flukes) that resemble fish fins due to convergent evolution, still have physiology based on their terrestrial ancestry, most notably their air-breathing respiration via lungs (instead of aquatic respiration via gills) and excretion of nitrogenous waste as urea or uric acid (instead of ammonia like most fish). Nearly all extant aquatic tetrapods are secondarily aquatic, with only larval amphibians (tadpoles) being primarily aquatic with gills, and only some species of paedomorphic mole salamanders (most notably the fully aquatic axolotl) retain the gill-based physiology into adulthood.

Acanthodii or acanthodians is an extinct class of gnathostomes (jawed fishes). They are currently considered to represent a paraphyletic grade of various fish lineages basal to extant Chondrichthyes, which includes living sharks, rays, and chimaeras. Acanthodians possess a mosaic of features shared with both osteichthyans (bony fish) and chondrichthyans (cartilaginous fish). In general body shape, they were similar to modern sharks, but their epidermis was covered with tiny rhomboid platelets like the scales of holosteians (gars, bowfins).

The popular name "spiny sharks" is because they were superficially shark-shaped, with a streamlined body, paired fins, a strongly upturned tail, and stout, largely immovable bony spines supporting all the fins except the tail—hence, "spiny sharks". However, acanthodians are not true sharks; their close relation to modern cartilaginous fish can lead them to be considered "stem-sharks". Acanthodians had a cartilaginous skeleton, but their fins had a wide, bony base and were reinforced on their anterior margin with a dentine spine. As a result, fossilized spines and scales are often all that remains of these fishes in ancient sedimentary rocks. The earliest acanthodians were marine, but during the Devonian, freshwater species became predominant.

Nekton or necton (from the Ancient Greek: νηκτόν, romanized: nekton, lit. 'to swim') is any aquatic organism that can actively and persistently propel itself through a water column (i.e. swim) without touching the bottom. Nekton generally have powerful tails and appendages (e.g. fins, pleopods, flippers or jets) that make them strong enough swimmers to counter ocean currents, and have mechanisms for sufficient lift and/or buoyancy to prevent sinking. Examples of extant nekton include most fish (especially pelagic fish like tuna and sharks), marine mammals (cetaceans, sirenia and pinnipeds) and reptiles (specifically sea turtles), penguins, coleoid cephalopods (squids and cuttlefish) and several species of decapod crustaceans (specifically prawns, shrimp and krill).

The term was proposed by German biologist Ernst Haeckel to differentiate between the active swimmers in a body of water, and the plankton that are passively carried along by the current. As a guideline, nektonic organisms have a high Reynolds number (greater than 1000) and planktonic organisms a low one (less than 10). Some organisms begin their life cycle as planktonic eggs and larvae, and transition to nektonic juveniles and adults later in life. This may make distinction difficult when attempting to classify certain plankton-to-nekton species as one or the other. For this reason, some biologists avoid using this term.

A fin is a thin appendage or component attached to a larger body or structure. Fins typically function as foils that produce lift or thrust, or provide the ability to steer or stabilize motion while traveling in water, air, or other fluids. Fins are also used to increase surface areas for heat transfer purposes, or simply as ornamentation.

Fins first evolved on fish as a means of locomotion. Fish fins are used to generate thrust and control the subsequent motion. Fish and other aquatic animals, such as cetaceans, actively propel and steer themselves with pectoral and tail fins. As they swim, they use other fins, such as dorsal and anal fins, to achieve stability and refine their maneuvering.

Actinopterygii (/ˌæktɪnɒptəˈrɪdʒiaɪ/ ; from Ancient Greek ἀκτίς (aktís) 'ray, beam' and πτέρυξ (ptérux) 'wing, fins'), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that constitute nearly 99% of the over 30,000 living species of fish. The vast majority of extant actinopterygian species are teleosts, and by species count they dominate the subphylum Vertebrata, comprising over 50% of all living vertebrates. They are the most abundant nektonic aquatic animals and are ubiquitous throughout freshwater, brackish and marine environments from the deep sea to subterranean waters to the highest mountain streams. Extant species can range in size from Paedocypris, at 8 mm (0.3 in), to the giant sunfish, at 2,700 kg (6,000 lb), and the giant oarfish, at 8 m (26 ft) (or possibly 11 m (36 ft)). The largest ever known ray-finned fish, the extinct Leedsichthys from the Jurassic, is estimated to have grown to 16.5 m (54 ft).

Ray-finned fish are so called because of their lightly built fins made of webbings of skin supported by radially extended thin bony spines called lepidotrichia, as opposed to the bulkier, fleshy fins of the sister clade Sarcopterygii (lobe-finned fish). Resembling folding fans, the actinopterygian fins can easily change shape, orientation and wetted area, providing superior thrust-to-weight ratios per movement compared to sarcopterygian and chondrichthyian fins. The fin rays attach directly to the proximal or basal skeletal elements, the radials, which represent the articulation between these fins and the internal skeleton (e.g., pelvic and pectoral girdles).

Fish go through various life stages between fertilization and adulthood. The life of fish start as spawned eggs which hatch into immotile larvae. These larval hatchlings are not yet capable of feeding themselves and carry a yolk sac which provides stored nutrition. Before the yolk sac completely disappears, the young fish must mature enough to be able to forage independently. When they have developed to the point where they are capable of feeding by themselves, the fish are called fry. When, in addition, they have developed scales and working fins, the transition to a juvenile fish is complete and it is called a fingerling, so called as they are typically about the size of human fingers. The juvenile stage lasts until the fish is fully grown, sexually mature and interacting with other adult fish.

The appendicular skeleton is the portion of the vertebrate endoskeleton consisting of the bones, cartilages and ligaments that support the paired appendages (fins, flippers or limbs). In most terrestrial vertebrates (except snakes, legless lizards and caecillians), the appendicular skeleton and the associated skeletal muscles are the predominant locomotive structures.

There are 126 bones in the human appendicular skeleton, includes the skeletal elements within the shoulder and pelvic girdles, upper and lower limbs, and hands and feet. These bones have shared ancestry (are homologous) to those in the forelimbs and hindlimbs of all other tetrapods, which are in turn homologous to the pectoral and pelvic fins in fish.