Eocene in the context of Thanetian

The Great Sphinx of Giza is a limestone statue of a reclining sphinx, a mythical creature with the head of a human and the body of a lion.

The monument was sculpted from the limestone bedrock of the Eocene-aged Mokattam Formation and faces east on the Giza Plateau, on the west bank of the Nile in Giza, Egypt. The oldest known monumental sculpture in Egypt, the Sphinx is part of the Memphite Necropolis and is a UNESCO World Heritage Site.

The Mokattam Formation is a Middle Eocene-aged geological formation in northern Egypt. Consisting of nummulitic marine limestone outcropping across the Mokattam Hills, it has been extensively quarried from Ancient Egypt to the present day, and represents the source material for most of the famous archeological sites of Greater Cairo, most notably the Giza pyramid complex, the Great Sphinx, and much of Historic Cairo.

The age of the formation is thought to span from the late Lutetian to the Bartonian, depending on the member (Building Stone or Giushi). However, some authors treat the Giushi Member as its own geologic formation, which would restrict the Mokattam to just the Lutetian.

Penguins are a group of flightless, semi-aquatic, sea birds which live almost exclusively in the Southern Hemisphere. Only one species, the Galápagos penguin, lives at, and slightly north of, the equator. Highly adapted for life in the ocean water, penguins have countershaded dark and white plumage and flippers for swimming. Most penguins feed on krill, fish, squid and other forms of sea life which they catch with their bills and swallow whole while swimming. A penguin has a spiny tongue and powerful jaws to grip slippery prey.

They spend about half of their lives on land and the other half in the sea. The largest living species is the emperor penguin (Aptenodytes forsteri): on average, adults are about 1.1 m (3 ft 7 in) tall and weigh 35 kg (77 lb). The smallest penguin species is the little blue penguin (Eudyptula minor), also known as the fairy penguin, which stands around 30–33 cm (12–13 in) tall and weighs 1.2–1.3 kg (2.6–2.9 lb). Today, larger penguins generally inhabit colder regions, and smaller penguins inhabit regions with temperate or tropical climates. Some prehistoric penguin species were enormous: as tall or heavy as an adult human. There was a great diversity of species in subantarctic regions, and at least one giant species in a region around 2,000 km south of the equator 35 mya, during the Late Eocene, a climate decidedly warmer than today.

The Eocene–Oligocene extinction event, also called the Eocene-Oligocene transition (EOT) or Grande Coupure (French for "great cut"), is the transition between the end of the Eocene and the beginning of the Oligocene, an extinction event and faunal turnover occurring between 33.9 and 33.4 million years ago. It was marked by large-scale extinction and floral and faunal turnover, although it was relatively minor in comparison to the largest mass extinctions.

Guinea fowl (/ˈɡɪnifaʊl/) are birds of the family Numididae in the order Galliformes. They are endemic to Africa and rank among the oldest of the gallinaceous birds. Phylogenetically, they branched off from the core Galliformes after the Cracidae (chachalacas, guans, and curassows) and before the Odontophoridae (New World quail). An Eocene fossil lineage Telecrex has been associated with guinea fowl; Telecrex inhabited Mongolia, and may have given rise to the oldest of the true phasianids, such as blood pheasants and eared pheasants, which evolved into high-altitude, montane-adapted species with the rise of the Tibetan Plateau. While modern guinea fowl species are endemic to Africa, the helmeted guinea fowl has been introduced as a domesticated bird widely elsewhere.

Strepsirrhini or Strepsirhini (/ˌstrɛpsəˈraɪni/ ; STREP-sə-RY-nee) is a suborder of primates that includes the lemuriform primates, which consist of the lemurs of Madagascar, galagos ("bushbabies") and pottos from Africa, and the lorises from India and Southeast Asia. Collectively they are referred to as strepsirrhines.

Also belonging to the suborder are the extinct adapiform primates, which thrived during the Eocene in Europe, North America, and Asia, but disappeared from most of the Northern Hemisphere as the climate cooled. Adapiforms are sometimes referred to as being "lemur-like", although the diversity of both lemurs and adapiforms does not support this comparison.

Tarsiiformes /ˈtɑːrsi.ɪfɔːrmiːz/ are a group of primates that once ranged across Europe, northern Africa, Asia, and North America, but whose extant species are all found in the islands of Southeast Asia. Tarsiers (family Tarsiidae) are the only living members of the infraorder; other members of Tarsiidae include the extinct Tarsius eocaenus from the Eocene, and Tarsius thailandicus from the Miocene. Two extinct genera, Xanthorhysis and Afrotarsius, are considered to be close relatives of the living tarsiers, and are generally classified within Tarsiiformes, with the former grouped within family Tarsiidae, and the latter listed as incertae sedis (undefined). Omomyids are generally considered to be extinct relatives, or even ancestors, of the living tarsiers, and are often classified within Tarsiiformes.

Other fossil primates, including Microchoeridae, Carpolestidae, and Eosimiidae, have been included in this classification, although the fossil evidence is debated. Eosimiidae has also been classified under the infraorder Simiiformes (with monkeys and apes), and most experts now consider Eosimiidae to be stem simians. Likewise, Carpolestidae is often classified within the order Plesiadapiformes, a very close, extinct relative of primates.

The Vespidae are a large (nearly 5000 species), diverse, cosmopolitan family of wasps, including nearly all the known eusocial wasps (such as Polistes fuscatus, Vespa orientalis, and Vespula germanica) and many solitary wasps. Each social wasp colony includes a queen and a number of female workers with varying degrees of sterility relative to the queen. In temperate social species, colonies usually last only one year, dying at the onset of winter. New queens and males (drones) are produced towards the end of the summer, and after mating, the queens hibernate over winter in cracks or other sheltered locations. The nests of most species are constructed out of mud, but polistines and vespines use plant fibers, chewed to form a sort of paper (also true of some stenogastrines). Many species are pollen vectors contributing to the pollination of several plants, being potential or even effective pollinators, while others are notable predators of pest insect species, and a few species are invasive pests.

The subfamilies Polistinae and Vespinae are composed solely of eusocial species, while the Eumeninae, Euparagiinae, Gayellinae, Masarinae and Zethinae are all solitary with the exception of a few communal and several subsocial species. The Stenogastrinae are facultatively eusocial, considering nests may have one or several adult females; in cases where the nest is shared by multiple females (typically, a mother and her daughters) there is reproductive division of labor and cooperative brood care.

Sir Charles Lyell, 1st Baronet, FRS (14 November 1797 – 22 February 1875) was a Scottish geologist who demonstrated the power of known natural causes in explaining the earth's history. He is best known today for his association with Charles Darwin and as the author of Principles of Geology (1830–33), which presented to a wide public audience the idea that the earth was shaped by the same natural processes still in operation today, operating at similar intensities. The philosopher William Whewell dubbed this gradualistic view "uniformitarianism" and contrasted it with catastrophism, which had been championed by Georges Cuvier and was better accepted in Europe. The combination of evidence and eloquence in Principles convinced a wide range of readers of the significance of "deep time" for understanding the earth and environment.

Lyell's scientific contributions included a pioneering explanation of climate change, in which shifting boundaries between oceans and continents could be used to explain long-term variations in temperature and rainfall. Lyell also gave influential explanations of earthquakes and developed the theory of gradual "backed up-building" of volcanoes. In stratigraphy his division of the Tertiary period into the Pliocene, Miocene, and Eocene was highly influential. He incorrectly conjectured that icebergs were the impetus behind the transport of glacial erratics, and that silty loess deposits might have settled out of flood waters. His creation of a separate period for human history, entitled the 'Recent', is widely cited as providing the foundations for the modern discussion of the Anthropocene.

The Great Sphinx of Giza is a limestone statue of a reclining sphinx, a mythical creature with the head of a human and the body of a lion. The monument was sculpted from the limestone bedrock of the Eocene-aged Mokattam Formation and faces east on the Giza Plateau, on the west bank of the Nile in Giza, Egypt. The oldest known monumental sculpture in Egypt, the Sphinx is part of the Memphite Necropolis and is a UNESCO World Heritage Site.

Archaeological evidence suggests the Sphinx was created by Egyptians of the Old Kingdom during the reign of Khufu (c. 2590–2566 BC) or Khafre (c. 2558–2532 BC). Scholars and Egyptologists believe the face of the Sphinx was carved to represent either the pharaoh Khufu or one of his sons, pharaohs Djedefre and Khafre, but a consensus has not been reached and the person(s) in whose likeness the Sphinx was carved remains in dispute.

A nummulite is a large lenticular fossil, characterised by its numerous coils, subdivided by septa into chambers. They are the shells of the fossil and present-day marine protozoan Nummulites, a type of foraminiferan. Nummulites commonly vary in diameter from 13 to 50 mm (0.51 to 1.97 in) and are common in Eocene to Miocene marine rocks, particularly around southwest Asia and the Mediterranean in the area that once constituted the Tethys Ocean, such as Eocene limestones from Egypt or from Pakistan. Fossils up to six inches wide are found in the Middle Eocene rocks of Turkey. They are valuable as index fossils.

The ancient Egyptians used nummulite shells as coins and the pyramids were constructed using limestone that contained nummulites. It is not surprising then that the name Nummulites is a diminutive form of the Latin nummulus 'little coin', a reference to their shape.

The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between 48.07 and 41.03 Ma. The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Middle Eocene Subepoch.

The Bartonian is, in the International Commission on Stratigraphy's (ICS) geologic time scale, a stage or age in the middle of the Eocene Epoch or Series. The Bartonian Age spans the time between 41.03 and 37.71 Ma. It is preceded by the Lutetian and is followed by the Priabonian Age.

Azolla (common called mosquito fern, water fern, and fairy moss) is a genus of seven species of aquatic ferns in the family Salviniaceae. They are extremely reduced in form and specialized, having a significantly different appearance to other ferns and more resembling some mosses or even duckweeds. Azolla filiculoides is one of two fern species for which a reference genome has been published. It is believed that this genus grew so prolifically during the Eocene (and thus absorbed such a large amount of carbon) that it triggered a global cooling event that has lasted to the present.

Azolla may establish as an invasive plant in areas where it is not native. In such a situation, it can alter aquatic ecosystems and biodiversity substantially by exhausting oxygen and covering water surface making underwater plants unable to photosynthesise.

Pteridospermatophyta, also called pteridosperms or seed ferns, are a polyphyletic grouping of extinct seed-producing plants. The earliest fossil evidence for plants of this type are the lyginopterids of late Devonian age. They flourished particularly during the Carboniferous and Permian periods. Pteridosperms declined during the Mesozoic Era and had mostly disappeared by the end of the Cretaceous Period, though Komlopteris seem to have survived into Eocene times, based on fossil finds in Tasmania.

With regard to the enduring utility of this division, many palaeobotanists still use the pteridosperm grouping in an informal sense to refer to the seed plants that are not angiosperms, coniferoids (conifers or cordaites), ginkgophytes (ginkgos or czekanowskiales), cycadophytes (cycads or bennettites), or gnetophytes. This is particularly useful for extinct seed plant groups whose systematic relationships remain speculative, as they can be classified as pteridosperms with no invalid implications being made as to their systematic affinities. Also, from a purely curatorial perspective the term pteridosperms is a useful shorthand for describing the fern-like fronds that were probably produced by seed plants, which are commonly found in many Palaeozoic and Mesozoic fossil floras.

Palaeotherium is an extinct genus of equoid that lived in Europe and possibly the Middle East from the Middle Eocene to the Early Oligocene. It is the type genus of the Palaeotheriidae, a group exclusive to the Palaeogene that was closest in relation to the Equidae, which contains horses plus their closest relatives and ancestors. Fossils of Palaeotherium were first described in 1782 by the French naturalist Robert de Lamanon and then closely studied by another French naturalist, Georges Cuvier, after 1798. Cuvier erected the genus in 1804 and recognized multiple species based on overall fossil sizes and forms. As one of the first fossil genera to be recognized with official taxonomic authority, it is recognized as an important milestone within the field of palaeontology. The research by early naturalists on Palaeotherium contributed to the developing ideas of evolution, extinction, and succession and demonstrating the morphological diversity of different species within one genus.

Since Cuvier's descriptions, many other naturalists from Europe and the Americas recognized many species of Palaeotherium, some valid, some reclassified to different genera afterward, and others being eventually rendered invalid. The German palaeontologist Jens Lorenz Franzen modernized its taxonomy due to his recognition of many subspecies as part of his dissertation in 1968, which were subsequently accepted by other palaeontologists. Today, there are sixteen known species recognized, many of which have multiple subspecies. In 1992, the French palaeontologist Jean-Albert Remy recognized two subgenera that most species are classified to based on cranial anatomies: the specialized Palaeotherium and the more generalized Franzenitherium.

An oak is a hardwood tree or shrub in the genus Quercus of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisphere, with some 500 species, both deciduous and evergreen. Fossil oaks date back to the Middle Eocene. Molecular phylogeny shows that the genus is divided into Old World and New World clades, but many oak species hybridise freely, making the genus's history difficult to resolve.

Ecologically, oaks are keystone species in habitats from Mediterranean semi-desert to subtropical rainforest. They live in association with many kinds of fungi including truffles. Oaks support more than 950 species of caterpillar, many kinds of gall wasp which form distinctive galls (roundish woody lumps such as the oak apple), and a large number of pests and diseases. Oak leaves and acorns contain enough tannin to be toxic to cattle, but pigs are able to digest them safely. Oak timber is strong and hard, and has found many uses in construction and furniture-making. The bark was traditionally used for tanning leather. Wine barrels are made of oak; these are used for aging alcoholic beverages such as sherry and whisky, giving them a range of flavours, colours, and aromas. The spongy bark of the cork oak is used to make traditional wine bottle corks. Almost a third of oak species are threatened with extinction due to climate change, invasive pests, and habitat loss.

The Oligocene is a geologic epoch of the Paleogene Period that extends from about 33.9 million to 23 million years before the present (33.9±0.1 to 23.04±0.05 Ma). As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from Ancient Greek ὀλίγος (olígos), meaning "few", and καινός (kainós), meaning "new", and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period.

The Oligocene is often considered an important time of transition, a link between the archaic world of the tropical Eocene and the more modern ecosystems of the Miocene. Major changes during the Oligocene included a global expansion of grasslands, and a regression of tropical broad leaf forests to the equatorial belt.

The Paleogene Period (IPA: /ˈpeɪli.ədʒiːn, -li.oʊ-, ˈpæli-/ PAY-lee-ə-jeen, -⁠lee-oh-, PAL-ee-; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period 66 Ma (million years ago) to the beginning of the Neogene Period 23.04 Ma. It is the first period of the Cenozoic Era, the tenth period of the Phanerozoic and is divided into the Paleocene, Eocene, and Oligocene epochs. The earlier term Tertiary Period was used to define the time now covered by the Paleogene Period and subsequent Neogene Period; despite no longer being recognized as a formal stratigraphic term, "Tertiary" still sometimes remains in informal use. Paleogene is often abbreviated "Pg", although the United States Geological Survey uses the abbreviation "Pe" for the Paleogene on the Survey's geologic maps.

Much of the world's modern vertebrate diversity originated in a rapid surge of diversification in the early Paleogene, as survivors of the Cretaceous–Paleogene extinction event took advantage of empty ecological niches left behind by the extinction of the non-avian dinosaurs, pterosaurs, marine reptiles, and primitive fish groups. Mammals continued to diversify from relatively small, simple forms into a highly diverse group ranging from small-bodied forms to very large ones, radiating into multiple orders and colonizing the air and marine ecosystems by the Eocene. Birds, the only surviving group of dinosaurs, quickly diversified from the very few neognath and paleognath clades that survived the extinction event, also radiating into multiple orders, colonizing different ecosystems and achieving an extreme level of morphological diversity. Percomorph fish, the most diverse group of vertebrates today, first appeared near the end of the Cretaceous but saw a very rapid radiation into their modern order and family-level diversity during the Paleogene, achieving a diverse array of morphologies.