Crustose in the context of "Photobiont"

A lichen (/ˈlaɪkən/ LIE-kən, UK also /ˈlɪtʃən/ LI-chən) is a hybrid colony of algae or cyanobacteria living symbiotically among filaments of multiple fungus species, along with bacteria embedded in the cortex or "skin", in a mutualistic relationship. Lichens are the lifeform that first brought the term symbiosis (as Symbiotismus) into biological context.

Lichens have since been recognized as important actors in nutrient cycling and producers which many higher trophic feeders feed on, such as reindeer, gastropods, nematodes, mites, and springtails. Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not plants. They may have tiny, leafless branches (fruticose) or flat, leaf-like structures (foliose); they may grow crust-like, adhering tightly to a surface (substrate) like a thick coat of paint (crustose), have a powder-like appearance (leprose), or feature other growth forms.

Coralline algae are red algae in the order Corallinales, characterized by a thallus containing calcareous deposits within its cell walls, giving it hardness. The colors of these algae are typically some hue of pink, or another shade of red, but some species can be purple, yellow, blue, white, or gray-green. Typically, these algae grow in a crustose manner (encrusting rocks and other hardscape); in the intertidal zone of rocky shorelines, and within coral reefs, these algae appear as an abundance of colorful patches on rock surfaces. Unattached specimens (maerl, rhodoliths) may form relatively smooth compact balls, or forming warty to fruticose thalli.

The red algae belong to the division Rhodophyta, within which the coralline algae form the order Corallinales. There are over 1600 described species of nongeniculate coralline algae. The corallines are presently grouped into two families on the basis of their reproductive structures. Most are marine, though one species lives in freshwater; Pneophyllum cetinaensis.

Graphidales is an order of lichen-forming fungi in the class Lecanoromycetes. It contains 6 families, about 81 genera and about 2,228 species. Family Graphidaceae are the largest crustose family within Graphidales order comprising more than 2000 species, which are widely distributed in tropical and subtropical regions of the world.

Rhodoliths (from Greek for red rocks) are colorful, unattached calcareous nodules, composed of crustose, benthic marine red algae that resemble coral. Rhodolith beds create biogenic habitat for diverse benthic communities. The rhodolithic growth habit has been attained by a number of unrelated coralline red algae, organisms that deposit calcium carbonate within their cell walls to form hard structures or nodules that resemble beds of coral.

Rhodoliths do not attach themselves to the rocky seabed. Rather, they roll like tumbleweeds along the seafloor until they become too large in size to be mobilised by the prevailing wave and current regime. They may then become incorporated into a semi-continuous algal mat or form an algal build-up. While corals are animals that are both autotrophic (photosynthesize via their symbionts) or heterotrophic (feeding on plankton), rhodoliths produce energy solely through photosynthesis (i.e. they can only grow and survive in the photic zone of the ocean).

In lichens, rhizines are multicellular root-like structures arising mainly from the lower surface. A lichen with rhizines is termed rhizinate, while a lichen lacking rhizines is termed erhizinate. Rhizines serve only to anchor the lichen to their substrate; they do not absorb nutrients, as plant roots do. Characteristics of the rhizines are used to identify lichens, for example, whether they are dense or sparse, uniformly distributed or clumped in specific areas, and straight or branched. Only foliose lichens may possess rhizines, not crustose or fruticose lichens, which lack a lower cortex.

Rhizohyphae are a type of attachment structure on some lichens. Rhizohyphae are more slender than rhizines and are one cell thick in diameter. Rhizohyphae often occur as a felt-like hyphal mass.

Caloplaca marina, the orange sea lichen, is a crustose, placodioid lichen. It has wide distribution, and can be found near the shore on rocks or walls. Calos in Greek means nice, placa in Greek is shield. Caloplaca therefore means ‘beautiful patches’.