Conodont in the context of "Jawless fish"

Marker horizons (also referred to as chronohorizons, key beds or marker beds) are stratigraphic units of the same age and of such distinctive composition and appearance, that, despite their presence in separate geographic locations, there is no doubt about their being of equivalent age (isochronous) and of common origin. Such clear markers facilitate the correlation of strata, and used in conjunction with fossil floral and faunal assemblages and paleomagnetism, permit the mapping of land masses and bodies of water throughout the history of the earth. They usually consist of a relatively thin layer of sedimentary rock that is readily recognized on the basis of either its distinct physical characteristics or fossil content and can be mapped over a very large geographic area. As a result, a key bed is useful for correlating sequences of sedimentary rocks over a large area. Typically, key beds were created as the result of either instantaneous events or (geologically speaking) very short episodes of the widespread deposition of a specific types of sediment. As the result, key beds often can be used for both mapping and correlating sedimentary rocks and dating them. Volcanic ash beds (tonsteins and bentonite beds) and impact spherule beds, and specific megaturbidites are types of key beds created by instantaneous events. The widespread accumulation of distinctive sediments over a geologically short period of time have created key beds in the form of peat beds, coal beds, shell beds, marine bands, black shales in cyclothems, and oil shales. A well-known example of a key bed is the global layer of iridium-rich impact ejecta that marks the Cretaceous–Paleogene boundary (K–T boundary).

Palynology, the study of fossil pollens and spores, routinely works out the stratigraphy of rocks by comparing pollen and spore assemblages with those of well-known layers—a tool frequently used by petroleum exploration companies in the search for new fields. The fossilised teeth or elements of conodonts are an equally useful tool.

Fish began evolving about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include Haikouichthys. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans.

The earliest jawed vertebrates probably developed during the late Ordovician period. They are first represented in the fossil record from the Silurian by two groups of fish: the armoured fish known as placoderms, which evolved from the ostracoderms; and the Acanthodii (or spiny sharks). The jawed fish that are still extant in modern days also appeared during the late Silurian: the Chondrichthyes (or cartilaginous fish) and the Osteichthyes (or bony fish). The bony fish evolved into two separate groups: the Actinopterygii (or ray-finned fish) and Sarcopterygii (which includes the lobe-finned fish).

The Late Ordovician mass extinction (LOME), sometimes known as the end-Ordovician mass extinction or the Ordovician–Silurian extinction, is the first of the "big five" major mass extinction events in Earth's history, occurring roughly 445 million years ago (Ma). It is often considered to be the second-largest-known extinction event just behind the end-Permian mass extinction, in terms of the percentage of genera that became extinct. Extinction was global during this interval, eliminating 49–60% of marine genera and nearly 85% of marine species. Under most tabulations, only the Permian–Triassic mass extinction exceeds the Late Ordovician mass extinction in biodiversity loss. The extinction event abruptly affected all major taxonomic groups and caused the disappearance of one third of all brachiopod and bryozoan families, as well as numerous groups of conodonts, trilobites, echinoderms, corals, bivalves and graptolites. Despite its taxonomic severity, the Late Ordovician mass extinction did not produce major changes to ecosystem structures compared to other mass extinctions, nor did it lead to any particular morphological innovations. Diversity gradually recovered to pre-extinction levels over the first 5 million years of the Silurian period.

The Late Ordovician mass extinction is traditionally considered to occur in two distinct pulses. The first pulse (interval), known as LOMEI-1, began at the boundary between the Katian and Hirnantian stages of the Late Ordovician epoch. This extinction pulse is typically attributed to the Late Ordovician glaciation, which abruptly expanded over Gondwana at the beginning of the Hirnantian and shifted the Earth from a greenhouse to icehouse climate. Cooling and a falling sea level brought on by the glaciation led to habitat loss for many organisms along the continental shelves, especially endemic taxa with restricted temperature tolerance and latitudinal range. During this extinction pulse, there were also several marked changes in biologically responsive carbon and oxygen isotopes. Marine life partially rediversified during the cold period and a new cold-water ecosystem, the "Hirnantia fauna", was established.

The Triassic–Jurassic (Tr-J) extinction event (TJME), often called the end-Triassic extinction, marks the boundary between the Triassic and Jurassic periods, 201.4 million years ago. It represents one of five major extinction events during the Phanerozoic, profoundly affecting life on land and in the oceans.

In the seas, about 23–34% of marine genera disappeared; corals, bivalves, brachiopods, bryozoans, and radiolarians suffered severe losses of diversity and conodonts were completely wiped out, while marine vertebrates, gastropods, and benthic foraminifera were relatively unaffected. On land, all archosauromorph reptiles other than crocodylomorphs, dinosaurs, and pterosaurs became extinct. Crocodylomorphs, dinosaurs, pterosaurs, and mammals were left largely untouched, allowing them to become the dominant land animals for the next 135 million years. Plants were likewise significantly affected by the crisis, with floral communities undergoing radical ecological restructuring across the extinction event.

Stage 10 of the Cambrian is the still-unnamed third and final stage of the Furongian series. It follows the Jiangshanian and precedes the Ordovician Tremadocian Stage. The proposed lower boundary is the first appearance of the trilobite Lotagnostus americanus around 491 million years ago, but other fossils are also being discussed (see below). The upper boundary is defined as the appearance of the conodont Iapetognathus fluctivagus which marks the beginning of the Tremadocian and is radiometrically dated as 486.85 million years ago.

The Dapingian is the third stage of the Ordovician system and the first stage of the Middle Ordovician series. It is preceded by the Floian and succeeded by the Darriwilian. The base of the Dapingian (and the top of the Floian) is defined as the first appearance of the conodont species Baltoniodus triangularis which happened about 471.3 million years ago. The Dapingian lasted for about 1.9 million years until about 469.4 million years ago.