Clasper in the context of "Jawed fish"

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⭐ Core Definition: Clasper

In biology, a clasper is a male anatomical structure found in some groups of animals, used in mating.

Male cartilaginous fish have claspers formed from the posterior portion of their pelvic fin which serve to channel semen into the female's cloaca during mating. The act of mating in some fish including sharks usually includes one of the claspers raised to allow water into the siphon through a specific orifice. The clasper is then inserted into the cloaca, where it opens like an umbrella to anchor its position. The siphon then begins to contract, expelling water and sperm. The claspers of many shark species have spines or hooks, which may hold them in place during copulation. Many male holocephalans, including living chimaeras, have cephalic claspers (tenacula) on their heads, which are thought to aid in holding the female during mating.

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👉 Clasper in the context of Jawed fish

Gnathostomata (/ˌnæθˈstɒmətə/; from Ancient Greek: γνάθος (gnathos) 'jaw' + στόμα (stoma) 'mouth') are jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all extant vertebrates, including all living bony fishes (both ray-finned and lobe-finned, including their terrestrial tetrapod relatives) and cartilaginous fishes, as well as extinct prehistoric fish such as placoderms and acanthodians. Most gnathostomes have retained ancestral traits like true teeth, a stomach, and paired appendages (pectoral and pelvic fins, limbs, wings, etc.). Other traits are elastin, horizontal semicircular canal of the inner ear, myelinated neurons, and an adaptive immune system which has discrete lymphoid organs (spleen and thymus) and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene.

It is now assumed that Gnathostomata evolved from ancestors that already possessed two pairs of paired fins. Until recently these ancestors, known as antiarchs, were thought to have lacked pectoral or pelvic fins. In addition to this, some placoderms were shown to have a third pair of paired appendages, that had been modified to claspers in males and pelvic basal plates in females — a pattern not seen in any other vertebrate group. The jawless Osteostraci are generally considered the closest sister taxon of Gnathostomata.

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Clasper in the context of Elasmobranch

Elasmobranchii (/ɪˌlæzməˈbræŋki/) is a subclass of Chondrichthyes or cartilaginous fish, including modern sharks (division Selachii), and batomorphs (division Batomorphi, including rays, skates, and sawfish). Members of this subclass are characterised by having five to seven pairs of gill slits opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil.

The definition of the clade is unclear with respect to fossil chondrichthyans. Some authors consider it as equivalent to Neoselachii (the crown group clade including modern sharks, rays, and all other descendants of their last common ancestor). Other authors use the name Elasmobranchii for a broader branch-based group of all chondrichthyans more closely related to modern sharks and rays than to Holocephali (the clade containing chimaeras and their extinct relatives). Important extinct groups of elasmobranchs sensu lato include the hybodonts (Order Hybodontiformes), xenacanths (order Xenacanthformes) and Ctenacanthiformes. These are also often referred to as "sharks" in reference to their similar anatomy and ecology to modern sharks.

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Clasper in the context of Cladoselache

Cladoselache ("branch shark") is an extinct genus of shark-like chondrichthyan (cartilaginous fish) from the Late Devonian (Famennian) of North America. It was similar in body shape to modern lamnid sharks (such as mako sharks and the great white shark), but was not closely related to lamnids or to any other modern (selachian) shark. As an early chondrichthyan, it had yet to evolve traits of modern sharks such as accelerated tooth replacement, a loose jaw suspension, enameloid teeth, and possibly claspers.

Some 20th century studies considered Cladoselache to be a basal (early-diverging) member of Elasmobranchii, the fork of cartilaginous fish which leads to modern sharks and rays. More recent studies have identified distinctive traits of the chondrocranium (cartilaginous braincase), dorsal fin spines, and pectoral fin bases. These newly identified features support a close relationship to symmoriiforms, a small group of bizarre chondrichthyans such as the bristle-spined Stethacanthus. Cladoselache and symmoriiforms may be more closely related to chimaeras (a modern group of unusual deep-sea fish) than to true sharks and rays.

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