Animal echolocation in the context of Doppler effect

Porpoises (/ˈpɔːrpəsɪz/) are small dolphin-like cetaceans classified under the family Phocoenidae. Although similar in appearance to dolphins, they are more closely related to narwhals and belugas. There are eight extant species of porpoise, all among the smallest of the toothed whales. Porpoises are distinguished from dolphins by their flattened, spade-shaped teeth distinct from the conical teeth of dolphins, and lack of a pronounced beak, although some dolphins (e.g. Hector's dolphin) also lack a pronounced beak. Porpoises, and other cetaceans, belong to the clade Cetartiodactyla with even-toed ungulates.

Porpoises range in size from the vaquita, at 1.4 metres (4 feet 7 inches) in length and 54 kilograms (119 pounds) in weight, to the Dall's porpoise, at 2.3 m (7 ft 7 in) and 220 kg (490 lb). Several species exhibit sexual dimorphism in that the females are larger than males. They have streamlined bodies and two limbs that are modified into flippers. Porpoises use echolocation as their primary sensory system. Some species are well adapted for diving to great depths. As all cetaceans, they have a layer of fat, or blubber, under the skin to keep them warm in cold water.

Oceanic dolphins or Delphinidae are a widely distributed family of dolphins that live in the sea. Close to forty extant species are recognised. They include several big species whose common names contain "whale" rather than "dolphin", such as the Globicephalinae (round-headed whales, which include the false killer whale and pilot whale). Delphinidae is a family within the superfamily Delphinoidea, which also includes the porpoises (Phocoenidae) and the Monodontidae (beluga whale and narwhal). River dolphins are relatives of the Delphinoidea.

Oceanic dolphins range in size from the 1.7-metre-long (5 ft 7 in) and 50-kilogram (110-pound) Maui's dolphin to the 9.4-metre (31 ft) and 10-metric-ton (11-short-ton) orca, the largest known dolphin. Several species exhibit sexual dimorphism; the males are larger than females. They have streamlined muscular bodies and two limbs that are modified into flippers. Though not quite as flexible as seals, some dolphins can travel at speeds 29 km/h (18 mph) for short distances. Most delphinids primarily eat fish, along with a smaller number of squid and small crustaceans, but some species specialise in eating squid, or, in the case of the orca, also eat marine mammals and birds. All, however, are purely carnivorous. They typically have between 100 and 200 teeth, although a few species have considerably fewer. Delphinids travel in large pods, which may number a thousand individuals in some species. Each pod forages over a range of tens to hundreds of square kilometres. Some pods have a loose social structure, with individuals frequently joining or leaving, but others seem to be more permanent, perhaps dominated by a male and a harem of females. Individuals communicate by sound, producing low-frequency whistles, and also produce high-frequency broadband clicks of 80–220 kHz, which are primarily used for echolocation. Gestation lasts from 10 to 18 months, and results in the birth of a single calf. Some species are well adapted for diving to great depths. They have a layer of fat, or blubber, under the skin to keep warm in the cold water.

Lazzaro Spallanzani (Italian pronunciation: [ˈladdzaro spallanˈtsaːni]; 12 January 1729 – 11 February 1799) was an Italian Catholic priest (for which he was nicknamed Abbé Spallanzani), biologist and physiologist who made important contributions to the experimental study of bodily functions, animal reproduction, and animal echolocation. His research on biogenesis paved the way for the downfall of the theory of spontaneous generation, a prevailing idea at the time that organisms develop from inanimate matters, though the final death blow to the idea was dealt by French scientist Louis Pasteur a century later.

His most important works were summed up in his book Expériences pour servir a l'histoire de la génération des animaux et des plantes (Experiences to Serve to the History of the Generation of Animals and Plants), published in 1785. Among his contributions were experimental demonstrations of fertilisation between ova and spermatozoa, and in vitro fertilisation.

Bats (order Chiroptera /kaɪˈrɒptərə/) are winged mammals; the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their long spread-out digits covered with a thin membrane or patagium. The smallest bat, and one of the smallest extant mammals, is Kitti's hog-nosed bat, which is 29–33 mm (1.1–1.3 in) in length, 150 mm (5.9 in) across the forearm and 2 g (0.071 oz) in mass. The largest bats are the flying foxes, with the giant golden-crowned flying fox (Acerodon jubatus) reaching a weight of 1.5 kg (3.3 lb) and having a wingspan of 1.6 m (5 ft 3 in).

The second largest order of mammals after rodents, bats comprise about 20% of all classified mammal species worldwide, with at least 1,500 known species. These were traditionally divided into two suborders: the largely fruit-eating megabats, and the echolocating microbats. But more recent evidence has supported dividing the order into Yinpterochiroptera and Yangochiroptera, with megabats as members of the former along with several species of microbats. Many bats are insectivores, and most of the rest are frugivores (fruit-eaters) or nectarivores (nectar-eaters). A few species feed on animals other than insects; for example, the vampire bats are haematophagous (feeding on blood). Most bats are nocturnal, and many roost in caves or other refuges; it is uncertain whether bats have these behaviours to escape predators. Bats are distributed globally in all except the coldest regions. They are important in their ecosystems for pollinating flowers and dispersing seeds as well as controlling insect populations.

The cetacean family Monodontidae comprises two living whale species, the narwhal and the beluga whale and at least four extinct species, known from the fossil record. Beluga and narwhal are native to coastal regions and pack ice around the Arctic Ocean. Both species are relatively small whales, 3–5 m (9.8–16.4 ft) in length, with a forehead melon, and a short or absent snout. Premaxillary teeth are absent. They do not have a true dorsal fin, but do have a narrow ridge running along the back, which is much more pronounced in the narwhal. They are highly vocal animals, communicating with a wide range of sounds. Like other whales, they also use echolocation to navigate. Belugas can be found in the far north of the Atlantic and Pacific Oceans; the distribution of narwhals is restricted to the Arctic and Atlantic Oceans.

Monodontids have a wide-ranging carnivorous diet, feeding on fish, molluscs, and small crustaceans. They have reduced teeth, with the beluga having numerous simple teeth, and the narwhal having only two teeth, one of which forms the tusks in males. Gestation lasts 14–15 months in both species, and almost always results in a single calf. The young are not weaned for a full two years, and do not reach sexual maturity until they are five to eight years of age. Family groups travel as part of herds, or 'pods', which may contain several hundred individuals.

River dolphins are a polyphyletic group of fully aquatic mammals that reside exclusively in freshwater or brackish water. They are an informal grouping of dolphins, which itself is a paraphyletic group within the infraorder Cetacea. Extant river dolphins are placed in two superfamilies, Platanistoidea and Inioidea. They comprise the families Platanistidae (the South Asian dolphins), the possibly extinct Lipotidae (Yangtze River dolphin), Iniidae (the Amazonian dolphins) and Pontoporiidae. There are five extant species of river dolphins. River dolphins, alongside other cetaceans, belong to the clade Artiodactyla, with even-toed ungulates, and their closest living relatives the hippopotamuses, from which they diverged about 40 million years ago. Specific types of dolphins can be pink.

River dolphins are relatively small compared to other dolphins, having evolved to survive in warm, shallow water and strong river currents. They range in size from the 5-foot (1.5 m) long South Asian river dolphin to the 8-foot (2.4 m) and 220-pound (100 kg) Amazon river dolphin. Several species exhibit sexual dimorphism, in that the females are larger than the males. They have streamlined bodies and two limbs that are modified into flippers. River dolphins use their conical-shaped teeth and long beaks to capture fast-moving prey in murky water. They have well-developed hearing that is adapted for both air and water; they do not really rely on vision since their eyes are very small and the water they swim in is usually very muddy. Instead, they tend to rely on echolocation when hunting and navigating. These species are well-adapted to living in warm, shallow waters, and, unlike other cetaceans, have little to no blubber.

The beluga whale (/bəˈluːɡə, bɪ-, bɛˈlʊɡə/; Delphinapterus leucas) is an Arctic and sub-Arctic cetacean. It is one of two living members of the family Monodontidae, along with the narwhal, and the only member of the genus Delphinapterus. It is also known as the white whale, as it is the only cetacean to regularly occur with this colour; the sea canary, due to its high-pitched calls; and the melonhead, though that more commonly refers to the melon-headed whale, which is an oceanic dolphin.

The beluga is adapted to life in the Arctic, with anatomical and physiological characteristics that differentiate it from other cetaceans. Amongst these are its all-white colour and the absence of a dorsal fin, which allows it to swim under ice with ease. It possesses a distinctive protuberance at the front of its head which houses an echolocation organ called the melon, which in this species is large and deformable. The beluga's body size is between that of a dolphin and a true whale, with males growing up to 5.5 m (18 ft) long and weighing up to 1,600 kg (3,530 lb). This whale has a stocky body. Like many cetaceans, a large percentage of its weight is blubber (subcutaneous fat). Its sense of hearing is highly developed and its echolocation allows it to move about and find breathing holes under sheet ice.

Horseshoe bats are bats in the family Rhinolophidae. In addition to the single living genus, Rhinolophus, which has about 106 species, the extinct genus Palaeonycteris has been recognized. Horseshoe bats are closely related to the Old World leaf-nosed bats, family Hipposideridae, which have sometimes been included in Rhinolophidae. The horseshoe bats are divided into six subgenera and many species groups. The most recent common ancestor of all horseshoe bats lived 34–40 million years ago, though it is unclear where the geographic roots of the family are, and attempts to determine its biogeography have been indecisive. Their taxonomy is complex, as genetic evidence shows the likely existence of many cryptic species, as well as species recognized as distinct that may have little genetic divergence from previously recognized taxa. They are found in the Old World, mostly in tropical or subtropical areas, including Africa, Asia, Europe, and Oceania.

Horseshoe bats are considered small or medium-sized microbats, weighing 4–28 g (0.14–0.99 oz), with forearm lengths of 30–75 mm (1.2–3.0 in) and combined lengths of head and body of 35–110 mm (1.4–4.3 in). The fur, long and smooth in most species, can be reddish-brown, blackish, or bright orange-red. They get their common name from their large nose-leafs, which are shaped like horseshoes. The nose-leafs aid in echolocation; horseshoe bats have highly sophisticated echolocation, using constant frequency calls at high-duty cycles to detect prey in areas of high environmental clutters. They hunt insects and spiders, swooping down on prey from a perch, or gleaning from foliage. Little is known about their mating systems, but at least one species is monogamous, while another is polygynous. Gestation is approximately seven weeks and one offspring is produced at a time. A typical lifespan is six or seven years, but one greater horseshoe bat lived more than thirty years.

Microbats constitute the suborder Microchiroptera within the order Chiroptera (bats). Bats have long been differentiated into Megachiroptera (megabats) and Microchiroptera, based on their size, the use of echolocation by the Microchiroptera and other features; molecular evidence suggests a somewhat different subdivision, as the microbats have been shown to be a paraphyletic group.

South Asian river dolphins are toothed whales in the genus Platanista, which inhabit the waterways of the Indian subcontinent. They were historically considered to be one species (P. gangetica) with the Ganges river dolphin and the Indus river dolphin being subspecies (P. g. gangetica and P. g. minor respectively). Genetic and morphological evidence led to their being described as separate species in 2021. The Ganges and Indus river dolphins are estimated to have diverged 550,000 years ago. They are the only living members of the family Platanistidae and the superfamily Platanistoidea. Fossils of ancient relatives date to the late Oligocene.

South Asian river dolphins are small but stocky cetaceans with long snouts or rostra, broad flippers, and small dorsal fins. They have several unusual features. Living in murky river waters, they have eyes that are tiny and lensless; the dolphins rely instead on echolocation for navigation. The skull has large crests over the melon, which help direct their echolocation signals. These dolphins prey mainly on fish and shrimp and hunt them throughout the water column. They are active through the day and are sighted in small groups. Both species are listed as endangered by the IUCN Red List of mammals. Major threats include dams, barrages, fishing nets, and both chemical and acoustic pollution.

The melon is a mass of adipose tissue found in the foreheads of all toothed whales. It focuses and modulates the animal's vocalizations and acts as a sound lens. It is thus a key organ involved in communication and echolocation.

Spermaceti /spərməˈsiːtiˌ -ˈsɛ-/ (see also: sperm oil) is a waxy substance found in the head cavities of the sperm whale (and, in smaller quantities, in the oils of other whales). Spermaceti is created in the spermaceti organ inside the whale's head. This organ may contain as much as 1,900 litres (500 US gal) of spermaceti. It has been extracted by whalers since the 17th century for human use in cosmetics, textiles, and candles.

Theories for the spermaceti organ's biological function suggest that it may control buoyancy, act as a focusing apparatus for the whale's sense of echolocation, or possibly both. Concrete evidence supports both theories. The buoyancy theory holds that the sperm whale is capable of heating the spermaceti, lowering its density and thus allowing the whale to float; for the whale to sink again, it must take water into its blowhole, which cools the spermaceti into a denser solid. This claim has been called into question by recent research that indicates a lack of biological structures to support this heat exchange, and the fact that the change in density is too small to be meaningful until the organ grows to a huge size. Measurement of the proportion of wax esters retained by a harvested sperm whale accurately described the age and future life expectancy of a given individual. The level of wax esters in the spermaceti organ increases with the age of the whale: 38–51% in calves, 58–87% in adult females, and 71–94% in adult males.

Bats (order Chiroptera /kaɪˈrɒptərə/) are winged mammals; the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their long spread-out digits covered with a thin membrane or patagium. The smallest bat, and one of the smallest extant mammals, is Kitti's hog-nosed bat, which is 29–33 mm (1.1–1.3 in) in length, 150 mm (5.9 in) across the forearm and 2 g (0.071 oz) in mass. The largest bats are the flying foxes, with the giant golden-crowned flying fox (Acerodon jubatus) reaching a weight of 1.5 kg (3.3 lb) and having a wingspan of 1.6 m (5 ft 3 in).

The second largest order of mammals after rodents, bats comprise about 20% of all classified mammal species worldwide, with at least 1,500 known species. These were traditionally divided into two suborders: the largely fruit-eating megabats and the echolocating microbats. But more recent evidence has supported dividing the order into Yinpterochiroptera and Yangochiroptera, with megabats as members of the former along with several species of microbats. Many bats are insectivores, and most of the rest are frugivores (fruit-eaters) or nectarivores (nectar-eaters). A few species feed on animals other than insects; for example, the vampire bats are haematophagous (feeding on blood). Most bats are nocturnal, and many roost in caves or other refuges; it is uncertain whether bats have these behaviours to escape predators. Bats are distributed globally in all except the coldest regions. They are important in their ecosystems for pollinating flowers and dispersing seeds as well as controlling insect populations.

The New World leaf-nosed bats (Phyllostomidae) are bats (order Chiroptera) found from southern North America to South America, specifically from the Southwest United States to northern Argentina. Both the scientific and common names derive from their often large, lance-shaped noses, though this is greatly reduced in some of the nectar- and pollen-feeders. Because these bats echolocate nasally, this "nose-leaf" is thought to serve some role in modifying and directing the echolocation call. Similar nose leaves are found in some other groups of bats, most notably the Old World leaf-nosed bats.

They are the most ecologically diverse bat family; members of this family have evolved to use food groups as varied as fruit, nectar, pollen, insects, frogs, other bats, and small vertebrates, and in the case of the vampire bats, blood. Most species are insectivorous, but the phyllostomid bats also include true predatory species and frugivores (subfamily Stenodermatinae and Carolliinae). For example, the spectral bat (Vampyrum spectrum), the largest bat in the Americas, eats vertebrate prey, including small, dove-sized birds.

Jacques-Yves Cousteau, AC (/kuːˈstoʊ/, also UK: /ˈkuːstoʊ/; French: [ʒak iv kusto]; 11 June 1910 – 25 June 1997) was a French naval officer, oceanographer, filmmaker and author. He co-invented the first successful open-circuit self-contained underwater breathing apparatus (SCUBA), called the Aqua-Lung, which assisted him in producing some of the first underwater documentaries.

Cousteau wrote many books describing his undersea explorations. In his first book, The Silent World: A Story of Undersea Discovery and Adventure, Cousteau surmised the existence of the echolocation abilities of porpoises. The book was adapted into an underwater documentary called The Silent World. Co-directed by Cousteau and Louis Malle, it was one of the first films to use underwater cinematography to document the ocean depths in color. The film won the 1956 Palme d'Or at the Cannes Film Festival and remained the only documentary to do so until 2004 (when Fahrenheit 9/11 received the award). It was also awarded the Academy Award for Best Documentary in 1957.