The Miaolingian is the third Series of the Cambrian Period, and was formally named in 2018. It lasted from about 506.5 to 497 million years ago and is divided in ascending order into 3 stages: the Wuliuan, Drumian, and Guzhangian. The Miaolingian is preceded by the unnamed Cambrian Series 2 and succeeded by the Furongian series.
It is named after the Miaoling Mountains in southeastern Guizhou Province, China.
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At 508 million years old (middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints.
The rock unit is a black shale and crops out at several localities near the town of Field in Yoho National Park and the Kicking Horse Pass. Another outcrop is in Kootenay National Park 42Â km to the south.
Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian (Miaolingian, Wuliuan) through the Lower Carboniferous (Mississippian). A possible early graptolite, Chaunograptus, is known from the Middle Cambrian. Recent analyses have favored the idea that the living pterobranch Rhabdopleura represents an extant graptolite which diverged from the rest of the group in the Cambrian.
Fossil graptolites and Rhabdopleura share a colony structure of interconnected zooids housed in organic tubes (theca) which have a basic structure of stacked half-rings (fuselli). Most extinct graptolites belong to two major orders: the bush-like sessile Dendroidea and the planktonic, free-floating Graptoloidea. These orders most likely evolved from encrusting pterobranchs similar to Rhabdopleura. Due to their widespread abundance, planktonic lifestyle, and well-traced evolutionary trends, graptoloids in particular are useful index fossils for the Ordovician and Silurian periods.