Vespidae in the context of Provespa

Sociality is the degree to which individuals in an animal population tend to associate in social groups (for which, the desire or inclination is known as gregariousness) and form cooperative societies.

Sociality is a survival response to evolutionary pressures. For example, when a mother wasp stays near her larvae in the nest, parasites are less likely to eat the larvae. Biologists suspect that pressures from parasites and other predators selected this behavior in wasps of the family Vespidae.

A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. The wasps do not constitute a clade, a complete natural group with a single ancestor, as bees and ants are deeply nested within the wasps, having evolved from wasp ancestors. Wasps that are members of the clade Aculeata can sting their prey.

The most commonly known wasps, such as yellowjackets and hornets, are in the family Vespidae and are eusocial, living together in a nest with an egg-laying queen and non-reproducing workers. Eusociality is favoured by the unusual haplodiploid system of sex determination in Hymenoptera, as it makes sisters exceptionally closely related to each other. However, the majority of wasp species are solitary, with each adult female living and breeding independently. Females typically have an ovipositor for laying eggs in or near a food source for the larvae, though in the Aculeata the ovipositor is often modified instead into a sting used for defense or prey capture.

Palaeovespa is an extinct genus of wasp in the Vespidae subfamily Vespinae. The genus currently contains eight species: five from the Priabonian stage Florissant Formation in Colorado, United States, two from the middle Eocene Baltic amber deposits of Europe, and one species from the late Paleocene of France.

The Oriental hornet (Vespa orientalis) is a social insect species of the family Vespidae. It can be found in Southwest Asia, Northeast Africa, the island of Madagascar (but no reports have been made of its presence on the island for many years), the Middle East, Central Asia, and parts of Southern Europe. Oriental hornets have also been found in a few isolated locations such as Mexico and Chile due to human introduction. The Oriental hornet lives in seasonal colonies consisting of caste system dominated by a queen. The hornet builds its nests underground and communicates using sound vibrations. The hornet has a yellow stripe on its cuticle (exoskeleton), which can absorb sunlight to generate a small electrical potential, and this might help supply energy for digging. The adult hornet eats nectar and fruits and scavenges for insects and animal proteins to feed to its young. Because they are scavengers, the hornets may also serve as a transmitter of disease following consumption of infected plants. The hornets are a primary pest to honey bees, attacking bee colonies to obtain honey and animal proteins. The sting of an Oriental hornet can be quite painful to humans and some humans are allergic to stings.

Vespula germanica, known colloquially as the European wasp, German wasp, or German yellowjacket, is a species of wasp found in much of the Northern Hemisphere, native to Europe, Northern Africa, and temperate Asia. It has spread and become well-established in many other places, including North America, South America (Argentina and Chile), Australia, South Africa, and New Zealand. German wasps are part of the family Vespidae and are sometimes mistakenly referred to as paper wasps because they build grey paper nests, although strictly speaking, paper wasps are part of the subfamily Polistinae. In North America, they are also known as yellowjackets.

The Polistinae is a subfamily of eusocial wasps belonging to the family Vespidae. They are closely related to the wasps (“yellowjackets” as they are called in North America) and true hornets of the subfamily Vespinae, containing four tribes. With about 1,100 species total, it is the second-most diverse subfamily within the Vespidae, and while most species are tropical or subtropical, they include some of the most frequently encountered large wasps in temperate regions.

The English common name for Polistinae is paper wasp. Many polistines, such as Polistes fuscatus, Polistes annularis, and Polistes exclamans, make their nests out of paper. Despite being called paper wasps, other wasps (including the wasps in the subfamily Vespinae) also build nests out of paper. Additionally, some epiponine wasps (such as Polybia emaciata) build their nests out of mud despite being Polistinae.

Potter wasps (or mason wasps), the Eumeninae, are a cosmopolitan wasp group currently considered a subfamily of Vespidae, but sometimes recognized in the past as a separate family, Eumenidae.

Mud dauber wasps, which also build their nests with mud, are in the families Sphecidae and Crabronidae and not discussed here.

The Euparagiinae are a small subfamily of rare wasps in the family Vespidae containing a single extant genus Euparagia. The group had a cosmopolitan distribution in past geological times extending back to the Early Cretaceous, but is now a geographically relict taxon known only from the desert regions of the Southwestern United States and northwestern Mexico.

This subfamily, with this same rank, has been previously grouped with the pollen wasps and treated together as the family "Masaridae". Now, the Euparagiinae are considered an independent subfamily and the sister group of the remainder of the Vespidae. Their wing venation is unique and differs from all other Vespidae; they also characteristically have a single small pale spot at the posterior edge of the mesonotum, and the femora and trochanters of the male front legs are modified in species-specific shapes.

Pollen wasps, the Masarinae, are unusual wasps that are typically treated as a subfamily of Vespidae, but have in the past sometimes been recognized as a separate family, "Masaridae", which also included the subfamilies Euparagiinae and Gayellinae. It is a small subfamily, unique among wasps in feeding their larvae exclusively with pollen and nectar, in a fashion quite similar to many solitary bees. Most species are black or brown, marked with strikingly contrasting patterns of yellow, white, or red (or combinations thereof). They are most diverse and abundant in the desert regions of southern Africa, but also occur in the deserts of North and South America. Some species of Pseudomasaris in California, such as Pseudomasaris vespoides, bear a remarkable resemblance to yellow jackets, but can be recognized by their strongly clubbed antennae, a characteristic feature of the subfamily. Males have greatly elongated antennae, but still ending in a strong club.

They carry pollen in their crops, and regurgitate it along with nectar when provisioning the cells of their nests, and they lay their eggs in the soupy mass before sealing the cell. The nests are often constructed of mud, or burrows in the ground, and these can have one to multiple individual cells. The nests are commonly located in concealed places, such as under rocks or in crevices.

The Stenogastrinae are a subfamily of social wasps included in the family Vespidae. They are sometimes called hover wasps owing to the particular hovering flight of some species. Their morphology and biology present interesting peculiarities.

Apoica flavissima is a paper wasp found primarily in South America. The species is distinguishable by its light coloring, unique single comb nests, and nocturnal nature. A notable feature of this species is the size dimorphism between queens and workers. Unlike most Vespidae wasps, Apocia flavissima queens are smaller than their worker counterparts which results in unique intraspecies relationships.

Ancistrocerus antilope is a species of wasp of the family Vespidae.