Tritylodontidae in the context of "Mammaliamorpha"

Cynodontia (from Ancient Greek κύων (kúōn) 'dog' and ὀδούς (odoús) 'tooth') is a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extinct ancestors and close relatives (Mammaliaformes), having evolved from advanced probainognathian cynodonts during the Late Triassic.

Non-mammalian cynodonts occupied a variety of ecological niches, both as carnivores and as herbivores. Following the emergence of mammals, most other cynodont lines went extinct, with the last known non-mammaliaform cynodont group, the Tritylodontidae, having its youngest records in the Early Cretaceous.

Epipubic bones are a pair of bones projecting forward from the pelvic bones of modern marsupials, monotremes and fossil mammals like multituberculates, and even basal eutherians (the ancestors of placentals, who lack them).They first occur in non-mammalian cynodonts such as tritylodontids, suggesting that they are a synapomorphy between them and Mammaliformes.

They were first described as early as 1698, but to date, their function(s) remain unresolved. Epipubic bones are often called marsupial bones because they support the mother's pouch in modern marsupials ("marsupium" is Latin for "pouch").

Mammaliaformes ("mammalian forms") is a clade of synapsid tetrapods that includes the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts during the Late Triassic. It is defined as the clade originating from the most recent common ancestor of Morganucodonta and the crown group mammals; the latter is the clade originating with the most recent common ancestor of extant Monotremata, Marsupialia and Placentalia. Besides Morganucodonta and the crown group mammals, Mammaliaformes also includes Docodonta and Hadrocodium.

Mammaliaformes is a term of phylogenetic nomenclature. In contrast, the assignment of organisms to class Mammalia has traditionally been founded on traits and, on this basis, Mammalia is slightly more inclusive than Mammaliaformes. In particular, trait-based taxonomy generally includes Adelobasileus and Sinoconodon in Mammalia, though they fall outside the Mammaliaformes definition. These genera are included in the broader clade Mammaliamorpha, defined phylogenetically as the clade originating with the last common ancestor of Tritylodontidae and the crown group mammals. This wider group includes some families that trait-based taxonomy does not include in Mammalia, in particular Tritylodontidae and Brasilodontidae.

Probainognathia is one of the two major subgroups of the clade Eucynodontia, the other being Cynognathia. The earliest forms were carnivorous and insectivorous, though some groups eventually also evolved herbivorous diets. The earliest and most basal probainognathian is the Middle Triassic (Anisian) aged Lumkuia, from South Africa, though probainognathians would not become prominent until the mid Norian stage of the Late Triassic. Three groups survived the extinction at the end of Triassic: Tritheledontidae and Tritylodontidae, which both survived until the Jurassic—the latter even into the Cretaceous (Montirictus and Xenocretosuchus)—and Mammaliaformes, which includes the mammals.