Tonian in the context of "Nerve cells"

The Neoproterozoic is the last of the three geologic eras of the Proterozoic eon, spanning from 1 billion to 538.8 million years ago, and is the last era of the Precambrian "supereon". It is preceded by the Mesoproterozoic era and succeeded by the Paleozoic era of the Phanerozoic eon, and is further subdivided into three periods, the Tonian, Cryogenian and Ediacaran.

One of the most severe glaciation events known in the geologic record occurred during the Cryogenian period of the Neoproterozoic, when global ice sheets may have reached the equator and created a "Snowball Earth" lasting about 100 million years. The earliest fossils of complex life are found in the Tonian period in the form of Otavia, a primitive sponge, and the earliest fossil evidence of metazoan radiation are found in the Ediacaran period, which included the namesaked Ediacaran biota as well as the oldest definitive cnidarians and bilaterians in the fossil record.

A neuron (American English), neurone (British English), or nerve cell, is an excitable cell that fires electric signals called action potentials across a neural network in the nervous system. They are located in the nervous system and help to receive and conduct impulses. Neurons communicate with other cells via synapses, which are specialized connections that commonly use minute amounts of chemical neurotransmitters to pass the electric signal from the presynaptic neuron to the target cell through the synaptic gap.

Neurons are the main components of nervous tissue in all animals except sponges and placozoans. Plants and fungi do not have nerve cells. Molecular evidence suggests that the ability to generate electric signals first appeared in evolution some 700 to 800 million years ago, during the Tonian period. Predecessors of neurons were the peptidergic secretory cells. They eventually gained new gene modules which enabled cells to create post-synaptic scaffolds and ion channels that generate fast electrical signals. The ability to generate electric signals was a key innovation in the evolution of the nervous system.

The Neoproterozoic Oxygenation Event (NOE), also called the Second Great Oxidation Event, was a geologic time interval between around 850 and 540 million years ago during the Neoproterozoic era, during which the oxygen concentration in Earth's atmosphere and oceans rose significantly. Taking place after the end to the Boring Billion, a euxinic period of extremely low atmospheric oxygen spanning from the Statherian period of the Paleoproterozoic era to the Tonian period of the Neoproterozoic era, the NOE was the second major increase in atmospheric and oceanic oxygen concentration on Earth, though it was not as prominent as the Great Oxidation Event (GOE) of the Neoarchean-Paleoproterozoic boundary. Unlike the GOE, it is unclear whether the NOE was a synchronous, global event or a series of asynchronous, regional oxygenation intervals with unrelated causes.

The Cryogenian (from Ancient Greek: κρύος, romanized: krýos, meaning "cold" and γένεσις, romanized: génesis, meaning "birth") is a geologic period that lasted from 720 to 635 million years ago. It is the second of the three periods of the Neoproterozoic era, preceded by the Tonian and followed by the Ediacaran.

The Cryogenian was a time of drastic climate changes. After the long environmental stability/stagnation during the Boring Billion, the Sturtian glaciation began at the beginning of Cryogenian, freezing the entire planet in a state of severe icehouse climate known as a snowball Earth. After 70 million years it ended, but was quickly followed by another global ice age, the Marinoan glaciation. There is controversy over whether these glaciations indeed covered the entire planet, or whether a band of open sea survived near the equator (i.e. "slushball Earth"), but the extreme climates with massive expanse of ice sheets blocking off sunlight would nevertheless have significantly hindered primary production in the shallow seas and caused major mass extinctions and biosphere turnovers.

Otavia antiqua is an ancient sponge-like multicellular organism found in the Otavi Group (the generic name being the namesake) in the Etosha National Park, Namibia. It is claimed to be the oldest animal fossil, being found in rock aged between 760 and 550 million years ago. The oldest Otavia fossils are from the Tonian period, before the Cryogenian glaciations, but the latest found were from the Nama Group rocks of the Ediacaran period.

The affinities of these fossils, along with other paleontological evidence for Precambrian sponges, are disputed.

The Boring Billion, otherwise known as the Mid Proterozoic and Earth's Middle Ages, is an informal geological time period between 1.8 and 0.8 billion years ago (Ga) during the middle Proterozoic eon spanning from the Statherian to the Tonian periods, characterized by more or less tectonic stability, climatic stasis and slow biological evolution. Although it is bordered by two different oxygenation events (the Great Oxygenation Event and Neoproterozoic Oxygenation Event) and two global glacial events (the Huronian and Cryogenian glaciations), the Boring Billion period itself actually had very low oxygen levels and no geological evidence of glaciations.

The oceans during the Boring Billion may have been oxygen-poor, nutrient-poor and sulfidic (euxinia), populated by mainly anoxygenic purple bacteria, a type of bacteriochlorophyll-based photosynthetic bacteria which uses hydrogen sulfide (H₂S) for carbon fixation instead of water and produces sulfur as a byproduct instead of oxygen. This is known as a Canfield ocean, and such composition may have caused the oceans to be colored black-and-milky-turquoise instead of blue or green as later. (By contrast, during the much earlier Purple Earth phase during the Archean, photosynthesis was performed mostly by archaeal colonies using retinal-based proton pumps that absorb green light, and the oceans would be magenta-purple.)

The Stenian Period (/ˈstiːni.ən/ STEE-nee-ən, from Ancient Greek: στενός, romanized: stenós, meaning "narrow") is the final geologic period in the Mesoproterozoic Era and lasted from 1200 Mya to 1000 Mya (million years ago). Instead of being based on stratigraphy, these dates are defined chronometrically. The name derives from narrow polymetamorphic belts formed over this period. It is preceded by the Ectasian Period and followed by the Neoproterozoic era and the Tonian period.

The supercontinent Rodinia assembled during the Stenian. It would last into the Tonian period before breaking up in the Cryogenian.