Synapsid in the context of "Temple (anatomy)"

The Permian (/ˈpɜːrmi.ən/ PUR-mee-ən) is a geologic period and stratigraphic system which spans 47 million years, from the end of the Carboniferous Period 298.9 Ma (million years ago) to the beginning of the Triassic Period 251.902 Ma. It is the sixth and last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia.

The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids (reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their amphibian ancestors.

A tetrapod (/ˈtɛtrəˌpɒd/; from Ancient Greek τετρα- (tetra-) 'four' and πούς (poús) 'foot') is any vertebrate animal of the clade Tetrapoda (/tɛˈtræpədə/). Tetrapods include all extant and extinct amphibians and amniotes, with the latter in turn evolving into two major clades, the sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (extinct "pelycosaurs", therapsids and all extant mammals, including humans). Hox gene mutations have resulted in some tetrapods becoming limbless (snakes, legless lizards, and caecilians) or two-limbed (cetaceans, sirenians, some lizards, kiwis, and the extinct moa and elephant birds). Nevertheless, they still qualify as tetrapods through their ancestry, and some retain a pair of vestigial spurs that are remnants of the hindlimbs.

Tetrapods evolved from a group of semiaquatic animals within the tetrapodomorphs which, in turn, evolved from ancient lobe-finned fish (sarcopterygians) around 390 million years ago in the Middle Devonian period. Early tetrapodomorphs were transitional between lobe-finned fishes and true four-limbed tetrapods, though most still fit the body plan expected of other lobe-finned fishes. The oldest fossils of four-limbed vertebrates (tetrapods in the broad sense of the word) are trackways from the Middle Devonian, and body fossils became common near the end of the Late Devonian, around 370–360 million years ago. These Devonian species all belonged to the tetrapod stem group, meaning that they did not belong to any modern tetrapod group.

The evolution of mammalian auditory ossicles was an evolutionary process that resulted in the formation of the mammalian middle ear, where the three middle ear bones or ossicles, namely the incus, malleus and stapes (a.k.a. "the anvil, hammer, and stirrup"), are a defining characteristic of mammals. The event is well-documented and important academically as a demonstration of transitional forms and exaptation, the re-purposing of existing structures during evolution.

The ossicles evolved from skull bones present in most tetrapods, including amphibians, sauropsids (which include extant reptiles and birds) and early synapsids (which include ancestors of mammals). The reptilian quadrate, articular and columella bones are homologs of the mammalian incus, malleus and stapes, respectively. In reptiles (and early synapsids by association), the eardrum is connected to the inner ear via a single bone, the columella, while the upper and lower jaws contain several bones not found in modern mammals. Over the course of mammalian evolution, one bone from the upper jaw (the quadrate) and one from the lower jaw (the articular) lost their function in the jaw articulation and migrated to form the middle ear. The shortened columella connected to these bones to form a kinematic chain of three ossicles, which serve to amplify air-sourced fine vibrations transmitted from the eardrum and facilitate more acute hearing in terrestrial environments.

Therapsida is a clade comprising a major group of eupelycosaurian synapsids that includes mammals and their ancestors and close relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, resulting in a more "standing" quadrupedal posture, as opposed to the lower sprawling posture of many reptiles and amphibians.

Therapsids evolved from earlier synapsids commonly called "pelycosaurs", specifically within the Sphenacodontia, more than 279.5 million years ago. They replaced the pelycosaurs as the dominant large land animals in the Guadalupian through to the Early Triassic. In the aftermath of the Permian–Triassic extinction event, therapsids declined in relative importance to the rapidly diversifying archosaurian sauropsids (pseudosuchians, dinosaurs and pterosaurs, etc.) during the Middle Triassic.

Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivores that typically bore a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian mass extinction that wiped out most other therapsids ca. 252 Mya. They rebounded at beginning of the following Triassic, but subsequently declined and died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 80-90 genera known, varying from rat-sized burrowers to elephant-sized browsers.

A piscivore (/ˈpɪsɪvɔːr/) is a carnivorous animal that primarily eats fish. Fish were the diet of early tetrapod evolution (via water-bound amphibians during the Devonian period); insectivory came next; then in time, the more terrestrially adapted reptiles and synapsids evolved herbivory.

Almost all predatory fish (most sharks, tuna, billfishes, pikes etc.) are obligated piscivores. Some non-piscine aquatic animals, such as whales, sea lions, and crocodilians, are not completely piscivorous; often also preying on invertebrates, marine mammals, waterbirds and even wading land animals in addition to fish, while others, such as the bulldog bat and gharial, are strictly dependent on fish for food. Some creatures, including cnidarians, octopuses, squid, cetaceans, spiders, grizzly bears, jaguars, wolves, snakes, turtles and sea gulls, may have fish as significant if not dominant portions of their diets. Humans can live on fish-based diets, as can their carnivorous domesticated pets such as dogs and cats.