Squamata in the context of "Scale (zoology)"

Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and amniotic development. Living traditional reptiles comprise four orders: Testudines, Crocodilia, Squamata, and Rhynchocephalia. About 12,000 living species of reptiles are listed in the Reptile Database. The study of the traditional reptile orders, customarily in combination with the study of modern amphibians, is called herpetology.

Reptiles have been subject to several conflicting taxonomic definitions. In evolutionary taxonomy, reptiles are gathered together under the class Reptilia (/rɛpˈtɪliə/ rep-TIL-ee-ə), which corresponds to common usage. Modern cladistic taxonomy regards that group as paraphyletic, since genetic and paleontological evidence has determined that crocodilians are more closely related to birds (class Aves), members of Dinosauria, than to other living reptiles, and thus birds are nested among reptiles from a phylogenetic perspective. Many cladistic systems therefore redefine Reptilia as a clade (monophyletic group) including birds, though the precise definition of this clade varies between authors. A similar concept is clade Sauropsida, which refers to all amniotes more closely related to modern reptiles than to mammals.

Snakes are elongated limbless reptiles of the suborder Serpentes (/sɜːrˈpɛntiːz/). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have skulls with several more joints than their lizard ancestors and relatives, enabling them to swallow prey much larger than their heads (cranial kinesis). To accommodate their narrow bodies, snakes' paired organs (such as kidneys) appear one in front of the other instead of side by side, and most only have one functional lung. Some species retain a pelvic girdle with a pair of vestigial claws on either side of the cloaca. Lizards have independently evolved elongate bodies without limbs or with greatly reduced limbs at least twenty-five times via convergent evolution, leading to many lineages of legless lizards. These resemble snakes, but several common groups of legless lizards have eyelids and external ears, which snakes lack, although this rule is not universal (see Amphisbaenia, Dibamidae, and Pygopodidae).

Living snakes are found on every continent except Antarctica, and on most smaller land masses; exceptions include some large islands, such as Ireland, Iceland, Greenland, and the islands of New Zealand, as well as many small islands of the Atlantic and central Pacific oceans. Additionally, sea snakes are widespread throughout the Indian and Pacific oceans. Around thirty families are currently recognized, comprising about 520 genera and about more than 4,170 species. They range in size from the tiny, 10.4 cm-long (4.1 in) Barbados threadsnake to the reticulated python of 6.95 meters (22.8 ft) in length. The fossil species Titanoboa cerrejonensis was 12.8 meters (42 ft) long. Snakes are thought to have evolved from either burrowing or aquatic lizards, perhaps during the Jurassic period, with the earliest known fossils dating to between 143 and 167 Ma ago. The diversity of modern snakes appeared during the Paleocene epoch (c. 66 to 56 Ma ago, after the Cretaceous–Paleogene extinction event). The oldest preserved descriptions of snakes can be found in the Brooklyn Papyrus.

Rhynchocephalia (/ˌrɪŋkoʊsɪˈfeɪliə/; lit. 'beak-heads') is an order of lizard-like reptiles that includes only one living species, the tuatara (Sphenodon punctatus) of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a speciose group with high morphological and ecological diversity. The oldest record of the group is dated to the Middle Triassic around 244 million years ago, and they had achieved global distribution by the Early Jurassic. Most rhynchocephalians belong to the suborder Sphenodontia ('wedge-teeth'). Their closest living relatives are lizards and snakes in the order Squamata, with the two orders being grouped together in the superorder Lepidosauria.

Rhynchocephalians are distinguished from squamates by a number of traits, including the retention of rib-like gastralia bones in the belly, as well as most rhynchocephalians having acrodont teeth that are fused to the crests of the jaws (the latter also found among a small number of modern lizard groups like agamids).

Mosasaurus (/ˌmoʊzəˈsɔːrəs/; "lizard of the Meuse River") is the type genus (defining example) of the mosasaurs, an extinct group of aquatic squamate reptiles. It lived from about 82 to 66 million years ago during the Campanian and Maastrichtian stages of the Late Cretaceous. The genus was one of the first Mesozoic marine reptiles known to science—the first fossils of Mosasaurus were found as skulls in a chalk quarry near the Dutch city of Maastricht in the late 18th century, and were initially thought to be crocodiles or whales. One skull discovered around 1780 was famously nicknamed the "great animal of Maastricht". In 1808, naturalist Georges Cuvier concluded that it belonged to a giant marine lizard with similarities to monitor lizards but otherwise unlike any known living animal. This concept was revolutionary at the time and helped support the then-developing ideas of extinction. Cuvier did not designate a scientific name for the animal; this was done by William Daniel Conybeare in 1822 when he named it Mosasaurus in reference to its origin in fossil deposits near the Meuse River. The exact affinities of Mosasaurus as a squamate remain controversial, and scientists continue to debate whether its closest living relatives are monitor lizards or snakes.

The largest species, M. hoffmannii, is estimated to measure up to 12 meters (39 ft) in maximum length, making it one of the largest mosasaurs. The skull of Mosasaurus had robust jaws and strong muscles capable of powerful bites using dozens of large teeth adapted for cutting prey. Its four limbs were shaped into paddles to steer the animal underwater. Its tail was long and ended in a downward bend and a paddle-like fluke. Mosasaurus possessed excellent vision to compensate for its poor sense of smell, and a high metabolic rate suggesting it was endothermic ("warm-blooded"), an adaptation in squamates only found in mosasaurs. There is considerable morphological variability across the currently-recognized species in Mosasaurus—from the robustly-built M. hoffmannii to the slender and serpentine M. lemonnieri—but an unclear diagnosis (description of distinguishing features) of the type species M. hoffmannii led to a historically problematic classification. As a result, more than fifty species have been attributed to the genus in the past. A redescription of the type specimen in 2017 helped resolve the taxonomy issue and confirmed at least five species to be within the genus. Another five species still nominally classified within Mosasaurus are planned to be reassessed.

Mosasaurs (from Latin Mosa meaning the 'Meuse', and Greek σαύρος sauros meaning 'lizard') are an extinct group of large aquatic reptiles within the family Mosasauridae that lived during the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. They belong to the order Squamata, which includes lizards and snakes.

During the last 20 million years of the Cretaceous period (Turonian–Maastrichtian ages), with the extinction of the ichthyosaurs and pliosaurs, mosasaurids became the dominant marine predators. They themselves became extinct as a result of the K-Pg event at the end of the Cretaceous period, about 66 million years ago.

Lizard is the common name used for all squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The grouping is paraphyletic as some lizards are more closely related to snakes than they are to other lizards. Lizards range in size from chameleons and geckos a few centimeters long to the 3-meter-long Komodo dragon.

Most lizards are quadrupedal, running with a strong side-to-side motion. Some lineages (known as "legless lizards") have secondarily lost their legs, and have long snake-like bodies. Some lizards, such as the forest-dwelling Draco, are able to glide. They are often territorial, the males fighting off other males and signalling, often with bright colours, to attract mates and to intimidate rivals. Lizards are mainly carnivorous, often being sit-and-wait predators; many smaller species eat insects, while the Komodo eats mammals as big as water buffalo.

Legless lizard may refer to any of several groups of lizards that have independently lost limbs or reduced them to the point of being of no use in locomotion. It is the common name for the family Pygopodidae. These lizards are often distinguishable from snakes on the basis of one or more of the following characteristics:

• possessing eyelids
• possessing external ear openings
• lack of broad belly scales
• notched rather than forked tongue
• having two lungs of roughly equal size (snakes have one short and one very long lung)
• having a very long tail (while snakes have a long body and short tail).

Every stage of reduction of the shoulder girdle —including complete loss— occurs among limbless squamates, but the pelvic girdle is never completely lost regardless of the degree of limb reduction or loss. At least the ilium is retained in limbless lizards and most basal snakes.