Root in the context of Stem tuber

A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor (anoxic) processes taking place, especially in the soils. Wetlands form a transitional zone between waterbodies and dry lands, and are different from other terrestrial or aquatic ecosystems due to their vegetation's roots having adapted to oxygen-poor waterlogged soils. They are considered among the most biologically diverse of all ecosystems, serving as habitats to a wide range of aquatic and semi-aquatic plants and animals, with often improved water quality due to plant removal of excess nutrients such as nitrates and phosphorus.

Wetlands exist on every continent, except Antarctica. The water in wetlands is either freshwater, brackish or saltwater. The main types of wetland are defined based on the dominant plants and the source of the water. For example, marshes are wetlands dominated by emergent herbaceous vegetation such as reeds, cattails and sedges. Swamps are dominated by woody vegetation such as trees and shrubs (although reed swamps in Europe are dominated by reeds, not trees). Mangrove forest are wetlands with mangroves and halophytic woody plants that have evolved to tolerate salty water.

The Silurian-Devonian Terrestrial Revolution, also known as the Devonian Plant Explosion (DePE) and the Devonian explosion, was a period of rapid colonization, diversification and radiation of land plants (particularly vascular plants) and fungi (especially dikaryans) on dry lands that occurred 428 to 359 million years ago (Mya) during the Silurian and Devonian periods, with the most critical phase occurring during the Late Silurian and Early Devonian.

This diversification of terrestrial photosynthetic florae had vast impacts on the biotic composition of the Earth's surface, especially upon the Earth's atmosphere by oxygenation and carbon fixation. Their roots also eroded into the rocks, creating a layer of water-holding and mineral/organic matter-rich soil on top of Earth's crust known as the pedosphere, and significantly altering the chemistry of Earth's lithosphere and hydrosphere. The floral activities following the Silurian-Devonian plant revolution also exerted significant influences on changes in the water cycle and global climate, as well as driving the biosphere by creating diverse layers of vegetations that provide both sustenance and refuge for both upland and wetland habitats, paving the way for all terrestrial and aquatic biomes that would follow.

The Devonian (/dəˈvoʊni.ən, dɛ-/ də-VOH-nee-ən, deh-) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.62 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.86 Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied.

The first significant evolutionary radiation of life on land occurred during the Devonian, as free-sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants (pteridospermatophytes) appeared. This rapid evolution and colonization process, which had begun during the Silurian, is known as the Silurian-Devonian Terrestrial Revolution. The earliest land animals, predominantly arthropods such as myriapods, arachnids and hexapods, also became well-established early in this period, after beginning their colonization of land at least from the Ordovician Period.

Groundwater recharge or deep drainage or deep percolation is a hydrologic process, where water moves downward from surface water to groundwater. Recharge is the primary method through which water enters an aquifer. This process usually occurs in the vadose zone below plant roots and is often expressed as a flux to the water table surface. Groundwater recharge also encompasses water moving away from the water table farther into the saturated zone. Recharge occurs both naturally (through the water cycle) and through anthropogenic processes (i.e., "artificial groundwater recharge"), where rainwater and/or reclaimed water is routed to the subsurface.

The most common methods to estimate recharge rates are: chloride mass balance (CMB); soil physics methods; environmental and isotopic tracers; groundwater-level fluctuation methods; water balance (WB) methods (including groundwater models (GMs)); and the estimation of baseflow (BF) to rivers.

A seagrass meadow or seagrass bed is an underwater ecosystem formed by seagrasses. Seagrasses are marine (saltwater) plants found in shallow coastal waters and in the brackish waters of estuaries. Seagrasses are flowering plants with stems and long green, grass-like leaves. They produce seeds and pollen and have roots and rhizomes which anchor them in seafloor sand.

Seagrasses form dense underwater meadows which are among the most productive ecosystems in the world. They provide habitats and food for a diversity of marine life comparable to that of coral reefs. This includes invertebrates like shrimp and crabs, cod and flatfish, marine mammals and birds. They provide refuges for endangered species such as seahorses, turtles, and dugongs. They function as nursery habitats for shrimps, scallops and many commercial fish species. Seagrass meadows provide coastal storm protection by the way their leaves absorb energy from waves as they hit the coast. They keep coastal waters healthy by absorbing bacteria and nutrients, and slow the speed of climate change by sequestering carbon dioxide into the sediment of the ocean floor.

Herbs are a widely distributed and widespread group of plants, excluding vegetables, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal purposes, or for fragrances. Culinary use typically distinguishes herbs from spices. Herbs generally refers to the leafy green or flowering parts of a plant (either fresh or dried), while spices are usually dried and produced from other parts of the plant, including seeds, bark, roots and fruits.

Herbs have a variety of uses including culinary, medicinal, aromatic and in some cases, spiritual. General usage of the term "herb" differs between culinary herbs and medicinal herbs; in medicinal or spiritual use, any parts of the plant might be considered "herbs", including leaves, roots, flowers, seeds, root bark, inner bark (and cambium), resin and pericarp.

A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, engages in photosynthesis, stores nutrients, and produces new living tissue. The stem can also be called the culm, halm, haulm, stalk, or thyrsus.

The stem is normally divided into nodes and internodes:

Bark is the outermost layer of stems and roots of woody plants. Plants with bark include trees, woody vines, and shrubs. Bark refers to all the tissues outside the vascular cambium and is a nontechnical term. It overlays the wood and consists of the inner bark and the outer bark. The inner bark, which in older stems is living tissue, includes the innermost layer of the periderm. The outer bark on older stems includes the dead tissue on the surface of the stems, along with parts of the outermost periderm and all the tissues on the outer side of the periderm. The outer bark on trees which lies external to the living periderm is also called the rhytidome.

Products derived from bark include bark shingle siding and wall coverings, spices, and other flavorings, tanbark for tannin, resin, latex, medicines, poisons, various hallucinogenic chemicals, and cork. Bark has been used to make cloth, canoes, and ropes and used as a surface for paintings and map making. A number of plants are also grown for their attractive or interesting bark colorations and surface textures or their bark is used as landscape mulch.

In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips of stems and roots, causing them to elongate, and gives rise to primary tissue. Secondary growth occurs in most seed plants, but monocots usually lack secondary growth. If they do have secondary growth, it differs from the typical pattern of other seed plants.

The formation of secondary vascular tissues from the cambium is a characteristic feature of dicotyledons and gymnosperms. In certain monocots, the vascular tissues are also increased after the primary growth is completed but the cambium of these plants is of a different nature. In the living pteridophytes this feature is extremely rare, only occurring in Isoetes.

Division, in horticulture and gardening, is a method of asexual plant propagation, where the plant (usually an herbaceous perennial) is broken up into two or more parts. Each part has an intact root and crown. The technique is of ancient origin, and has long been used to propagate bulbs such as garlic and saffron. Another type of division is through a plant tissue culture. In this method the meristem (a type of plant tissue) is divided.

A plant cutting is a piece of a plant that is used in horticulture for vegetative (asexual) propagation. A piece of the stem or root of the source plant is placed in a suitable medium such as moist soil. If the conditions are suitable, the plant piece will begin to grow as a new plant independent of the parent, a process known as striking. A stem cutting produces new roots, and a root cutting produces new stems. Some plants can be grown from leaf pieces, called leaf cuttings, which produce both stems and roots. The scions used in grafting are also called cuttings.

Propagating plants from cuttings is an ancient form of cloning. There are several advantages of cuttings, mainly that the produced offspring are practically clones of their parent plants. If a plant has favorable traits, it can continue to pass down its advantageous genetic information to its offspring. This is especially economically advantageous as it allows commercial growers to clone a certain plant to ensure consistency throughout their crops.

Grafting or graftage is a horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion (/ˈsaɪən/) while the lower part is called the rootstock. The success of this joining requires that the vascular tissues grow together. The natural equivalent of this process is inosculation. The technique is most commonly used in asexual propagation of commercially grown plants for the horticultural and agricultural trades. The scion is typically joined to the rootstock at the soil line; however, top work grafting may occur far above this line, leaving an understock consisting of the lower part of the trunk and the root system.

In most cases, the stock or rootstock is selected for its roots and the scion is selected for its stems, leaves, flowers, or fruits. The scion contains the desired genes to be duplicated in future production by the grafted plant.

Woodchips are small- to medium-sized pieces of wood formed by cutting or chipping larger pieces of wood such as trees, branches, logging residues, stumps, roots, and wood waste.

Woodchips may be used as a biomass solid fuel and are raw material for producing wood pulp. They may also be used as an organic mulch in gardening, landscaping, and ecosystem restoration; in bioreactors for denitrification; and as a substrate for mushroom cultivation.

The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (land plants with vascular tissues such as xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from non-vascular plants (mosses, hornworts and liverworts) by having specialized transport bundles that conduct water and nutrients from and to the roots, as well as life cycles in which the branched sporophyte is the dominant phase.

Ferns have complex leaves called megaphylls that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns that produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns and ophioglossoid ferns.

In botany, succulent plants, also known as succulents, are plants with parts that are thickened, fleshy, and engorged, usually to retain water in arid climates or soil conditions.

Succulents may store water in various structures, such as leaves and stems. The water content of some succulent organs can get up to 90–95%, such as Glottiphyllum semicyllindricum and Mesembryanthemum barkleyii. Some definitions also include roots, thus geophytes that survive unfavorable periods by dying back to underground storage organs (caudex) may be regarded as succulents. The habitats of these water-preserving plants are often in areas with high temperatures and low rainfall, such as deserts, but succulents may be found even in alpine ecosystems growing in rocky or sandy soil. Succulents are characterized by their ability to thrive on limited water sources, such as mist and dew, which makes them equipped to survive in ecosystems that contain scarce water sources.

In plant morphology, thorns, spines, and prickles, and in general spinose structures (sometimes called spinose teeth or spinose apical processes), are hard, rigid extensions or modifications of leaves, roots, stems, or buds with sharp, stiff ends, and generally serve the same function: physically defending plants against herbivory.

Soil gases (soil atmosphere) are the gases found in the air space between soil components. The spaces between the solid soil particles, if they do not contain water, are filled with air. The primary soil gases are nitrogen, carbon dioxide and oxygen. Oxygen is critical because it allows for respiration of both plant roots and soil organisms. Other natural soil gases include nitric oxide, nitrous oxide, methane, and ammonia. Some environmental contaminants below ground produce gas which diffuses through the soil such as from landfill wastes, mining activities, and contamination by petroleum hydrocarbons which produce volatile organic compounds. The soil atmosphere is also made of a variety of volatile compounds emitted by soil organisms, as respiratory metabolites, allelopathic compounds or semiochemical signals used in within-species and between-species communication. Soil is a net emitter of greenhouse gases, in particular when and where permafrost is thawing and degassing under the influence of climate warming.

Gases fill soil pores in the soil structure as water drains or is removed from a soil pore by evaporation or root absorption. The network of pores within the soil aerates, or ventilates, the soil. This aeration network becomes blocked when water enters soil pores. Not only are both soil air and soil water very dynamic parts of soil, but both are often inversely related.

A staple food, food staple, or simply staple, is a food that is eaten often and in such quantities that it constitutes a dominant portion of a standard diet for an individual or a population group, supplying a large fraction of energy needs and generally forming a significant proportion of the intake of other nutrients as well. For humans, a staple food of a specific society may be eaten as often as every day or every meal, and most people live on a diet based on just a small variety of food staples. Specific staples vary from place to place, but typically are inexpensive or readily available foods that supply one or more of the macronutrients and micronutrients needed for survival and health: carbohydrates, proteins, fats, minerals and vitamins. Typical examples include grains (cereals and legumes), seeds, nuts and root vegetables (tubers and roots). Among them, cereals (rice, wheat, oat, maize, etc.), legumes (lentils and beans) and tubers (e.g. potato, taro and yam) account for about 90% of the world's food calorie intake.

Early agricultural civilizations valued the crop foods that they established as staples because, in addition to providing necessary nutrition, they generally are suitable for storage over long periods of time without decay. Such nonperishable foods are the only possible staples during seasons of shortage, such as dry seasons or cold temperate winters, against which times harvests have been stored. During seasons of surplus, wider choices of foods may be available.

In biology, being semiaquatic refers to various macro-organisms that live regularly in both aquatic and terrestrial environments. When referring to animals, the term describes those that actively spend part of their daily time in water (in which case they can also be called amphibious), or land animals that have spent at least one life stage (e.g. as eggs or larvae) in aquatic environments. When referring to plants, the term describes land plants whose roots have adapted well to tolerate regular, prolonged submersion in water, as well as emergent and (occasionally) floating-leaved aquatic plants that are only partially immersed in water.

Examples of semiaquatic animals and plants are given below.

Nymphaeaceae (/ˌnɪmfiˈeɪsi.iː, -ˌaɪ/) is a family of flowering plants, commonly called water lilies. They live as rhizomatous aquatic herbs in temperate and tropical climates around the world. The family contains five genera with about 70 known species. Water lilies are rooted in soil in bodies of water, with leaves and flowers floating on or rising from the surface. Leaves are oval and heart-shaped in Barclaya. Leaves are round, with a radial notch in Nymphaea and Nuphar, but fully circular in Victoria and Euryale.

Water lilies are a well-studied family of plants because their large flowers with multiple unspecialized parts were initially considered to represent the floral pattern of the earliest flowering plants. Later genetic studies confirmed their evolutionary position as basal angiosperms. Analyses of floral morphology and molecular characteristics and comparisons with a sister taxon, the family Cabombaceae, indicate, however, that the flowers of extant water lilies with the most floral parts are more derived than the genera with fewer floral parts. Genera with more floral parts, Nuphar, Nymphaea, Victoria, have a beetle pollination syndrome, while genera with fewer parts are pollinated by flies or bees, or are self- or wind-pollinated. Thus, the large number of relatively unspecialized floral organs in the Nymphaeaceae is not an ancestral condition for the clade.