Pterosaurs in the context of "Convergently evolved"

Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last common ancestor of those groups. The cladistic term for the same phenomenon is homoplasy. The recurrent evolution of flight is a classic example, as flying insects, birds, pterosaurs, and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are analogous, whereas homologous structures or traits have a common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions.

The opposite of convergent evolution is divergent evolution, where related species evolve different traits. Convergent evolution is similar to parallel evolution, which occurs when two independent species evolve in the same direction and thus independently acquire similar characteristics; for instance, gliding frogs have evolved in parallel from multiple types of tree frog.

The Triassic (/traɪˈæsɪk/; sometimes symbolized as 🝈) is a geologic period and a stratigraphic system that spans 50.5 million years from the end of the Permian Period 251.902 Ma (million years ago) to the beginning of the Jurassic Period 201.4 Ma. The Triassic Period is the first and shortest geologic period of the Mesozoic Era, and the seventh period of the Phanerozoic Eon. The start and the end of the Triassic Period featured major extinction events.

Chronologically, the Triassic Period is divided into three epochs: (i) the Early Triassic, (ii) the Middle Triassic, and (iii) the Late Triassic. The Triassic Period began after the Permian–Triassic extinction event that much reduced the biosphere of planet Earth. The fossil record of the Triassic Period presents three categories of organisms: (i) animals that survived the Permian–Triassic extinction event, (ii) new animals that briefly flourished in the Triassic biosphere, and (iii) new animals that evolved and dominated the Mesozoic Era. Reptiles, especially archosaurs, were the chief terrestrial vertebrates during this time. A specialized group of archosaurs, called dinosaurs, first appeared in the Late Triassic but did not become dominant until the succeeding Jurassic Period. Archosaurs that became dominant in this period were primarily pseudosuchians, relatives and ancestors of modern crocodilians, while some archosaurs specialized in flight, the first time among vertebrates, becoming the pterosaurs. Therapsids, the dominant vertebrates of the preceding Permian period, saw a brief surge in diversification in the Triassic, with dicynodonts and cynodonts quickly becoming dominant, but they declined throughout the period with the majority becoming extinct by the end. However, the first stem-group mammals (mammaliamorphs), themselves a specialized subgroup of cynodonts, appeared during the Triassic and would survive the extinction event, allowing them to radiate during the Jurassic. Amphibians were primarily represented by the temnospondyls, giant aquatic predators that had survived the end-Permian extinction and saw a new burst of diversification in the Triassic, before going extinct by the end; however, early crown-group lissamphibians (including stem-group frogs, salamanders and caecilians) also became more common during the Triassic and survived the extinction event. The earliest known neopterygian fish, including early holosteans and teleosts, appeared near the beginning of the Triassic, and quickly diversified to become among the dominant groups of fish in both freshwater and marine habitats.

In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals. The other options are digitigrade, walking on the toes and fingers with the heel and wrist permanently raised, and unguligrade, walking on the nail or nails of the toes (the hoof) with the heel/wrist and the digits permanently raised. The leg of a plantigrade mammal includes the bones of the upper leg (femur/humerus) and lower leg (tibia and fibula/radius and ulna). The leg of a digitigrade mammal also includes the metatarsals/metacarpals, the bones that in a human compose the arch of the foot and the palm of the hand. The leg of an unguligrade mammal also includes the phalanges, the finger and toe bones.

Among extinct animals, most early mammals such as pantodonts were plantigrade. A plantigrade foot is the primitive condition for mammals; digitigrade and unguligrade locomotion evolved later. Among archosaurs, the pterosaurs were partially plantigrade and walked on the whole of the hind foot and the fingers of the hand-wing. Out of the plantigrade animals, only a few, such as humans, kangaroos and certain rodents, are obligate bipeds, while most others are functional bipeds.

Pseudosuchia (from Ancient Greek ψεύδος (pseúdos), meaning "false", and Σοῦχος (Soûkhos), meaning "Sobek") is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs", in contrast to the "bird-line archosaurs" or Avemetatarsalia. Despite Pseudosuchia meaning "false crocodiles", the name is a misnomer as true crocodilians are now defined as a subset of the group.

The clade Pseudosuchia is potentially equivalent to another term, Crurotarsi, even though the latter has a different, node-based definition: "all taxa the least inclusive clade containing Rutiodon carolinensis (Emmons, 1856), and Crocodylus niloticus (Laurenti, 1768)." Many paleontologists of the late 20th century took this proposal for granted, using Crurotarsi as the term for crocodilian ancestors. In 2011, a major revision of Triassic archosaur relations proposed that Rutiodon's group, Phytosauria, was not as closely related to other traditional "crurotarsans", at least compared to avemetatarsalians such as pterosaurs and dinosaurs. Under that interpretation, Crurotarsi would be a much broader clade than Pseudosuchia. Other recent studies continue to support a more traditional phylogeny with phytosaurs as an early branch of Pseudosuchia. If the traditional interpretation is maintained, Pseudosuchia and Crurotarsi are roughly equivalent categories.

The Niobrara Formation /ˌnaɪ.əˈbrærə/, also called the Niobrara Chalk, is a geologic formation in North America that was deposited between 87 and 82 million years ago during the Coniacian, Santonian, and Campanian stages of the Late Cretaceous. It is composed of two structural units, the Smoky Hill Chalk Member overlying the Fort Hays Limestone Member. The chalk formed from the accumulation of coccoliths from microorganisms living in what was once the Western Interior Seaway, an inland sea that divided the continent of North America during much of the Cretaceous. It underlies much of the Great Plains of the US and Canada. Evidence of vertebrate life is common throughout the formation and includes specimens of plesiosaurs, mosasaurs, pterosaurs, and several primitive aquatic birds. The type locality for the Niobrara Chalk is the Niobrara River in Knox County in northeastern Nebraska. The formation gives its name to the Niobrara cycle of the Western Interior Seaway.