Anomura (sometimes Anomala) is a group of decapod crustaceans, including hermit crabs and others. Several groups such as the king crabs and porcelain crabs within the Anomura have independently undergone carcinisation, acquiring the armoured crab body plan with a concealed tail. It is the sister group to the "true crabs" or Brachyura.
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Porcelain crab in the context of Decapod
The Decapoda or decapods, from Ancient Greek δεκάς (dekás), meaning "ten", and πούς (poús), meaning "foot", is a large order of crustaceans within the class Malacostraca, and includes crabs, lobsters, crayfish, shrimp, and prawns. Most decapods are scavengers. The order is estimated to contain nearly 15,000 extant species in around 2,700 genera, with around 3,300 fossil species. Nearly half of these species are crabs, with the shrimp (about 3,000 species) and Anomura including hermit crabs, king crabs, porcelain crabs, squat lobsters (about 2500 species) making up the bulk of the remainder. The earliest fossils of the group date to the Devonian.
Porcelain crab in the context of Brachyura
Brachyura (from Ancient Greek βραχύς (brakhús), meaning "short", and οὐρά (ourá), meaning "tail"), is an infraorder of decapod crustaceans comprising the true crabs. They typically have a very short projecting tail-like abdomen, usually hidden entirely under the thorax. Their exoskeleton is often thickened and hard. They generally have five pairs of legs, and they have "pincers" or "claws" on the ends of the frontmost pair, scientifically termed the chelae. They are present in all the world's oceans, in freshwater, and on land, often hiding themselves in small crevices or burrowing into sediment. True crabs are omnivores, feeding on a variety of food, including a significant proportion of algae, as well as detritus and other invertebrates. True crabs are widely consumed by humans as food, with over 1.5 million tonnes caught annually.
True crabs first appeared in the fossil record during the Jurassic period, around 200 million years ago, achieving great diversity by the Cretaceous period; around 7,000 extant species in 96 families are known. The true crabs are the largest of the crab decapod groups with carcinised body forms. Smaller groups include the porcelain crabs, king crabs, and hairy stone crabs which have independently evolved similar forms and lifestyles.
Porcelain crab in the context of Carcinisation
Carcinisation (American English: carcinization) is a form of convergent evolution in which non-crab crustaceans evolve a crab-like body plan. The term was introduced into evolutionary biology by Lancelot Alexander Borradaile, who described it in 1916 as "the many attempts of Nature to evolve a crab".