Middle Triassic in the context of Maculicorpus microbialis

The Triassic (/traɪˈæsɪk/; sometimes symbolized as 🝈) is a geologic period and a stratigraphic system that spans 50.5 million years from the end of the Permian Period 251.902 Ma (million years ago) to the beginning of the Jurassic Period 201.4 Ma. The Triassic Period is the first and shortest geologic period of the Mesozoic Era, and the seventh period of the Phanerozoic Eon. The start and the end of the Triassic Period featured major extinction events.

Chronologically, the Triassic Period is divided into three epochs: (i) the Early Triassic, (ii) the Middle Triassic, and (iii) the Late Triassic. The Triassic Period began after the Permian–Triassic extinction event that much reduced the biosphere of planet Earth. The fossil record of the Triassic Period presents three categories of organisms: (i) animals that survived the Permian–Triassic extinction event, (ii) new animals that briefly flourished in the Triassic biosphere, and (iii) new animals that evolved and dominated the Mesozoic Era. Reptiles, especially archosaurs, were the chief terrestrial vertebrates during this time. A specialized group of archosaurs, called dinosaurs, first appeared in the Late Triassic but did not become dominant until the succeeding Jurassic Period. Archosaurs that became dominant in this period were primarily pseudosuchians, relatives and ancestors of modern crocodilians, while some archosaurs specialized in flight, the first time among vertebrates, becoming the pterosaurs. Therapsids, the dominant vertebrates of the preceding Permian period, saw a brief surge in diversification in the Triassic, with dicynodonts and cynodonts quickly becoming dominant, but they declined throughout the period with the majority becoming extinct by the end. However, the first stem-group mammals (mammaliamorphs), themselves a specialized subgroup of cynodonts, appeared during the Triassic and would survive the extinction event, allowing them to radiate during the Jurassic. Amphibians were primarily represented by the temnospondyls, giant aquatic predators that had survived the end-Permian extinction and saw a new burst of diversification in the Triassic, before going extinct by the end; however, early crown-group lissamphibians (including stem-group frogs, salamanders and caecilians) also became more common during the Triassic and survived the extinction event. The earliest known neopterygian fish, including early holosteans and teleosts, appeared near the beginning of the Triassic, and quickly diversified to become among the dominant groups of fish in both freshwater and marine habitats.

Placozoa (/ˌplækəˈzoʊə/ PLAK-ə-ZOH-ə; lit. 'flat animals') is a phylum of free-living (non-parasitic) marine invertebrates. They are blob-like animals composed of aggregations of cells. Moving in water by ciliary motion, eating food by engulfment, reproducing by fission or budding, placozoans are described as "the simplest animals on Earth". Structural and molecular analyses have supported them as among the most basal animals, thus constituting a primitive metazoan phylum.

The first known placozoan, Trichoplax adhaerens, was discovered in 1883 by the German zoologist Franz Eilhard Schulze (1840–1921). Describing the uniqueness, another German, Karl Gottlieb Grell (1912–1994), erected a new phylum, Placozoa, for it in 1971. Remaining a monotypic phylum for over a century, new species began to be added since 2018. So far, three other extant species have been described, in two distinct classes: Uniplacotomia (Hoilungia hongkongensis in 2018 and Cladtertia collaboinventa in 2022) and Polyplacotomia (Polyplacotoma mediterranea, the most basal, in 2019). A single putative fossil species is known, the Middle Triassic Maculicorpus microbialis.

The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between 251.902 Ma and 246.7 Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a unit in chronostratigraphy. The Early Triassic is the oldest epoch of the Mesozoic Era. It is preceded by the Lopingian Epoch (late Permian, Paleozoic Era) and followed by the Middle Triassic Epoch. The Early Triassic is divided into the Induan and Olenekian ages. The Induan is subdivided into the Griesbachian and Dienerian ages and the Olenekian is subdivided into the Smithian and Spathian ages.

The Lower Triassic series is coeval with the Scythian Stage, which is today not included in the official timescales but can be found in older literature. In Europe, most of the Lower Triassic is composed of Buntsandstein, a lithostratigraphic unit of continental red beds.

The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between 237 Ma and 201.4 Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian ages.

Many of the first dinosaurs evolved during the Late Triassic, including Plateosaurus, Coelophysis, Herrerasaurus, and Eoraptor. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth.

Rhynchocephalia (/ˌrɪŋkoʊsɪˈfeɪliə/; lit. 'beak-heads') is an order of lizard-like reptiles that includes only one living species, the tuatara (Sphenodon punctatus) of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a speciose group with high morphological and ecological diversity. The oldest record of the group is dated to the Middle Triassic around 244 million years ago, and they had achieved global distribution by the Early Jurassic. Most rhynchocephalians belong to the suborder Sphenodontia ('wedge-teeth'). Their closest living relatives are lizards and snakes in the order Squamata, with the two orders being grouped together in the superorder Lepidosauria.

Rhynchocephalians are distinguished from squamates by a number of traits, including the retention of rib-like gastralia bones in the belly, as well as most rhynchocephalians having acrodont teeth that are fused to the crests of the jaws (the latter also found among a small number of modern lizard groups like agamids).

The Moenkopi Formation is a geological formation that is spread across the U.S. states of New Mexico, northern Arizona, Nevada, southeastern California, eastern Utah and western Colorado. This unit is considered to be a group in Arizona. Part of the Colorado Plateau and Basin and Range, this red sandstone was laid down in the Lower Triassic and possibly part of the Middle Triassic, around 240 million years ago.

In the geologic timescale, the Olenekian is an age in the Early Triassic epoch; in chronostratigraphy, it is a stage in the Lower Triassic series. It spans the time between 249.9 Ma and 246.7 Ma (million years ago). The Olenekian is sometimes divided into the Smithian and the Spathian subages or substages. The Olenekian follows the Induan and is followed by the Anisian (Middle Triassic).

The Olenekian saw the deposition of a large part of the Buntsandstein in Europe. The Olenekian is roughly coeval with the regional Yongningzhenian Stage used in China.

In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from 246.7 million years ago until 241.464 million years ago. The Anisian Age succeeds the Olenekian Age (part of the Lower Triassic Epoch) and precedes the Ladinian Age.

The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between 241.464 Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic).

The Ladinian is coeval with the Falangian regional stage used in China.

The Muschelkalk (German for "shell-bearing limestone", pronounced [ˈmʊʃl̩ˌkalk] ) is a sequence of sedimentary rock strata (a lithostratigraphic unit) in the geology of central and western Europe. It has a Middle Triassic (240 to 230 million years) age and forms the middle part of the three-part Germanic Trias (that gives the Triassic its name) lying above the older Buntsandstein and below the younger Keuper. The Muschelkalk ("mussel-chalk") consists of a sequence of limestone and dolomite beds.

In the past, the time span in which the Muschelkalk was deposited could also be called "Muschelkalk". In modern stratigraphy, however, the name only applies to the layers of rock.

The Keuper is a lithostratigraphic unit (a sequence of rock strata) in the subsurface of large parts of west and central Europe. The Keuper consists of dolomite, shales or claystones and evaporites that were deposited during the Middle and Late Triassic epochs (about 220 million years ago). The Keuper lies on top of the Muschelkalk and under the predominantly Lower Jurassic Lias or other Early Jurassic strata.

The Keuper together with the Muschelkalk and the Buntsandstein form the Germanic Trias Group, a characteristic sequence of rock strata that gave the Triassic its name.

Probainognathia is one of the two major subgroups of the clade Eucynodontia, the other being Cynognathia. The earliest forms were carnivorous and insectivorous, though some groups eventually also evolved herbivorous diets. The earliest and most basal probainognathian is the Middle Triassic (Anisian) aged Lumkuia, from South Africa, though probainognathians would not become prominent until the mid Norian stage of the Late Triassic. Three groups survived the extinction at the end of Triassic: Tritheledontidae and Tritylodontidae, which both survived until the Jurassic—the latter even into the Cretaceous (Montirictus and Xenocretosuchus)—and Mammaliaformes, which includes the mammals.

The tuatara (/tuːəˈtɑːrə/, Māori: [ˈtʉ.a.ta.ɾa]; Sphenodon punctatus) is a species of reptile endemic to New Zealand. Despite its close resemblance to lizards, it is the only extant member of a distinct lineage, the previously highly diverse order Rhynchocephalia. The name tuatara is derived from the Māori language and means "peaks on the back".

The single extant species of tuatara is the only surviving member of its order, which was highly diverse during the Mesozoic era. Rhynchocephalians first appeared in the fossil record during the Middle Triassic, around 244-241.5 million years ago, and reached worldwide distribution and peak diversity during the Jurassic, when they represented the world's dominant group of small reptiles. Rhynchocephalians declined during the Cretaceous, with their youngest records outside New Zealand dating to the Paleocene. Their closest living relatives are squamates (lizards and snakes). Tuatara are of interest for studying the evolution of reptiles.