The Middle Miocene is a sub-epoch of the Miocene epoch made up of two stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene, and followed by the Late Miocene.
The sub-epoch lasted from 15.97 ± 0.05 Ma (million years ago) to 11.608 ± 0.005 Ma. During this period, a sharp drop in global temperatures took place. This event is known as the Middle Miocene Climatic Transition.
Cainotheriidae is an extinct family of artiodactyls known from the Late Eocene to Middle Miocene of Europe. They are mostly found preserved in karstic deposits.
These animals were small in size, and generally did not exceed 15 centimetres (5.9 in) in height at the shoulders, ranging in size from those of rabbits to tragulids. For a long time, they were considered to have a similar lifestyle to hares and rabbits. The dentition was full and highly selenodont, i.e. the premolars and molars had curved and crescent-shaped cutting edges (as in today's ruminants). The skull was small, with a short snout and orbits closed posteriorly placed at the center of the skull.
Hucho is a genus of large piscivorous salmonid fish known as taimens (from Finnish taimen, 'trout', through Russian: тайме́нь, romanized: taĭménʹ), and is closely related to Pacific trout and lenoks (all belonging to the same tribe in the subfamily Salmoninae). Native to the cold rivers and other freshwater habitats in Eurasia, they are threatened by overfishing and habitat loss.
The earliest fossil remains of this genus are known from the Late Oligocene to middle Miocene of the Vitim Plateau in Russia. Younger remains are also known from the Late Miocene of Ukraine and the Late Pleistocene of Germany. Fossil specimens of a Hucho-like salmonid have been recovered from the Clarkia fossil beds and other localities from the late Neogene of western North America, suggesting they may have potentially inhabited North America too.
Equini is the only living tribe of the subfamily Equinae, which has lived worldwide (except Australia) since the Hemingfordian stage of the Middle Miocene (16–0 mya). It is considered to be a monophyletic clade.
The Astaracian age is a period of geologic time (16 to 11.6 Ma), equivalent with the Middle Miocene and used more specifically with European Land Mammal Ages. It precedes the Vallesian age and follows the Orleanian age. The Astaracian overlaps the Langhian and Serravallian ages.
During the Late Orleanian and Astaracian (17 to 10.7 Ma), oscillating sea levels resulted in a succession of palaeogeographic changes in the Eastern Mediterranean; the opening and closing of the Tethys seaway resulted in temporary land-bridges between Africa and Eurasia. Three short periods of faunal migrations between the continents can be distinguished:
The Vallesian age is a period of geologic time (11.6–9.0 Ma) within the Miocene used more specifically with European Land Mammal Ages. It precedes the Turolian age and follows the Astaracian age. The so-called Vallesian Crisis resulted in the extinction of several mammalian taxa characteristic of the Middle Miocene.
The term "Vallesian" was introduced by Catalan palaeontologist Miquel Crusafont in 1950 to mark the arrival of the equid Hipparion in Europe. The remaining European palaeofaunas, however, had been around since the Middle Miocene, including the moschid Micromeryx (a musk deer), the cervid Euprox, the suid Listriodon, and the felids Sansanosmilus and Pseudaelurus, and the Aragonian-Vallesian boundary does not represent a major shift in the European mammalian record. In contrast, the transition between Lower and Upper Vallesian corresponds to a major biotic crisis — the demise of most Aragonian artiodactyls, including the antelope Protragocerus, the bovid Miotragocerus, Listriodon, and the suids Hyotherium and Parachleusastochoerus. The crisis also affected rodents such as the family Eomyidae and most of the cricetids and glirids. They were replaced by species arriving from the east, Turolian in character: for example the suid Schizochoerus, the murid Progonomys, the bovids Tragoportax and Graecoryx, the hyaenid Adcrocuta, the felid Paramachairodus, and the suid Microstonyx.
The Clarendonian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 13,600,000 to 10,300,000 years BP, a period of 3.3 million years.
It is usually considered to overlap the Serravallian age of the Middle Miocene and the Tortonian age of the Late Miocene. The Clarendonian is preceded by the Barstovian and followed by the Hemphillian NALMA stages.
Danuvius guggenmosi is an extinct species of great ape that lived 11.6 million years ago during the Middle–Late Miocene in southern Germany. It is the sole member of the genus Danuvius. The area at this time was probably a woodland with a seasonal climate. A male specimen was estimated to have weighed about 31 kg (68 lb), and two females 17 and 19 kg (37 and 42 lb). Both genus and species were described in November 2019.
It is the first-discovered Late Miocene great ape with preserved long bones which could be used to reconstruct the limb anatomy and thus the locomotion of contemporary apes. Its discoverer, paleoanthropologist Madelaine Böhme, says Danuvius had adaptations for both hanging in trees (suspensory behavior) and walking on two legs (bipedalism)—whereas, among present-day great apes, humans are better adapted for the latter and the others the former. Danuvius thus had a method of locomotion unlike any previously known ape called "extended limb clambering", she says, walking directly along tree branches as well as using arms for suspending itself. The last common ancestor between humans and other apes possibly had a similar method of locomotion. However, paleoanthropologist Scott Williams and others say the fragmentary remains do not differ enough from other fossil apes to provide such a clue to the origins of bipedalism.