James City County is a county located in the Commonwealth of Virginia. As of the 2020 census, the population was 78,254. Although politically separate from the county, the county seat is the adjacent independent city of Williamsburg. Located on the Virginia Peninsula, James City County is included in the Virginia Beach-Norfolk-Newport News, VA-NC Metropolitan Statistical Area. It is often associated with Williamsburg, an independent city, and Jamestown which is within the county.
First settled by the English colonists in 1607 at Jamestown in the Virginia Colony, the county was formally created in 1634 as James City Shire by order of King Charles I. James City County is considered one of only five original shires of Virginia to still be extant today in essentially the same political form. The Jamestown 2007 celebration marked the 400th anniversary of the founding of Jamestown.