Hymenoptera in the context of "Beetles"

Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22,000 species have been classified. They are easily identified by their geniculate (elbowed) antennae and the distinctive node-like structure that forms their slender waists.

Ants form colonies that range in size from a few dozen individuals often living in small natural cavities to highly organised colonies that may occupy large territories with a sizeable nest (or nests) that consist of millions of individuals, in some cases they reach hundreds of millions of individuals in super colonies. Typical colonies consist of various castes of sterile, wingless females, most of which are workers (ergates), as well as soldiers (dinergates) and other specialised groups. Nearly all ant colonies also have some fertile males called "drones" and one or more fertile females called "queens" (gynes). The colonies are described as superorganisms because the ants appear to operate as a unified entity, collectively working together to support the colony.

Silk is a natural protein fiber, some forms of which can be woven into textiles. The protein fiber of silk is composed mainly of fibroin. It is most commonly produced by certain insect larvae to form cocoons. The best-known silk is obtained from the cocoons of the larvae of the mulberry silkworm Bombyx mori, which are reared in captivity (sericulture). The shimmery appearance of silk is due to the triangular prism-like structure of the silk fiber, which causes silk cloth to refract incoming light at different angles, thus producing different colors.

Harvested silk is produced by numerous insects; generally, only the silk of various moth caterpillars has been used for textile manufacturing. Research into other types of silk, which differ at the molecular level, has been conducted. Silk is produced primarily by the larvae of insects undergoing complete metamorphosis, but some insects, such as webspinners and raspy crickets, produce silk throughout their lives. Silk production also occurs in hymenoptera (bees, wasps, and ants), silverfish, caddisflies, mayflies, thrips, leafhoppers, beetles, lacewings, fleas, flies, and midges. Other types of arthropods also produce silk, most notably various arachnids, such as spiders.

A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. The wasps do not constitute a clade, a complete natural group with a single ancestor, as bees and ants are deeply nested within the wasps, having evolved from wasp ancestors. Wasps that are members of the clade Aculeata can sting their prey.

The most commonly known wasps, such as yellowjackets and hornets, are in the family Vespidae and are eusocial, living together in a nest with an egg-laying queen and non-reproducing workers. Eusociality is favoured by the unusual haplodiploid system of sex determination in Hymenoptera, as it makes sisters exceptionally closely related to each other. However, the majority of wasp species are solitary, with each adult female living and breeding independently. Females typically have an ovipositor for laying eggs in or near a food source for the larvae, though in the Aculeata the ovipositor is often modified instead into a sting used for defense or prey capture.

Eusociality (Greek εὖ eu 'good' and social) is the highest level of organization of sociality. It is defined by the following characteristics: cooperative brood care (including care of offspring from other individuals), overlapping generations within a colony of adults, and a division of labor into reproductive and non-reproductive groups. The division of labor creates specialized behavioral groups within an animal society, sometimes called castes. Eusociality is distinguished from all other social systems because individuals of at least one caste usually lose the ability to perform behaviors characteristic of individuals in another caste. Eusocial colonies can be viewed as superorganisms.

Eusociality has evolved among the insects, crustaceans, trematoda and mammals. It is most widespread in the Hymenoptera (ants, bees, and wasps) and in Isoptera (termites). A colony has caste differences: queens and reproductive males take the roles of the sole reproducers, while soldiers and workers work together to create and maintain a living situation favorable for the brood. Queens produce multiple queen pheromones to create and maintain the eusocial state in their colonies; they may also eat eggs laid by other females or exert dominance by fighting. There are two eusocial rodents: the naked mole-rat and the Damaraland mole-rat. Some shrimps, such as Synalpheus regalis, are eusocial. E. O. Wilson and others have claimed that humans have evolved a weak form of eusociality. It has been suggested that the colonial and epiphytic staghorn fern, too, may make use of a primitively eusocial division of labor.

Bees are winged insects that form a monophyletic clade Anthophila within the superfamily Apoidea of the order Hymenoptera, with over 20,000 known species in seven recognized families. Some species – including honey bees, bumblebees, and stingless bees – are social insects living in highly hierarchical colonies, while over 90% of bee species – including mason bees, carpenter bees, leafcutter bees, and sweat bees – are solitary. Members of the most well-known bee genus, Apis (i.e. honey bees), are known to construct hexagonally celled waxy nests called hives.

Unlike the closely related wasps and ants, who are carnivorous/omnivorous, bees are herbivores that specifically feed on nectar (nectarivory) and pollen (palynivory), the former primarily as a carbohydrate source for metabolic energy, and the latter primarily for protein and other nutrients for their larvae. They are found on every continent except Antarctica, and in every habitat on the planet that contains insect-pollinated flowering plants. The most common bees in the Northern Hemisphere are the Halictidae, or sweat bees, but they are small and often mistaken for wasps or flies. Bees range in size from tiny stingless bee species, whose workers are less than 2 millimeters (0.08 in) long, to the leafcutter bee Megachile pluto, the largest species of bee, whose females can attain a length of 39 millimeters (1.54 in). Vertebrate predators of bees include primates and birds such as bee-eaters; insect predators include beewolves and dragonflies.

A hyperparasite, also known as a metaparasite, is a parasite whose host is itself a parasite, often specifically a parasitoid. Hyperparasites are found mainly among the wasp-waisted Apocrita within the Hymenoptera, and in two other insect orders, the Diptera (true flies) and Coleoptera (beetles). Seventeen families in Hymenoptera and a few species of Diptera and Coleoptera are hyperparasitic. Hyperparasitism developed from primary parasitism, which evolved in the Jurassic period in the Hymenoptera. Hyperparasitism intrigues entomologists because of its multidisciplinary relationship to evolution, ecology, behavior, biological control, taxonomy, and mathematical models.

Biological control or biocontrol is a method of controlling pests, whether pest animals such as insects and mites, weeds, or pathogens affecting animals or plants by using other organisms. It relies on predation, parasitism, herbivory, or other natural mechanisms, but typically also involves an active human management role. It can be an important component of integrated pest management (IPM) programs.

There are three basic strategies for biological control: classical (importation), where a natural enemy of a pest is introduced in the hope of achieving control; inductive (augmentation), in which a large population of natural enemies are administered for quick pest control; and inoculative (conservation), in which measures are taken to maintain natural enemies through regular reestablishment.