Honey bee in the context of Southeast Asia

A natural language or ordinary language is any spoken language or signed language used organically in a human community, first emerging without conscious premeditation and subject to: replication across generations of people in the community, regional expansion or contraction, and gradual internal and structural changes. The vast majority of languages in the world are natural languages. As a category, natural language includes both standard dialects (ones with high social prestige) as well as nonstandard or vernacular dialects. Even an official language with a regulating academy such as Standard French, overseen by the Académie Française, is still classified as a natural language (e.g. in the field of natural language processing), as its prescriptive aspects do not make it regulated enough to be considered a constructed or controlled natural language. Linguists broadly consider writing to be a static visual representation of a particular natural language, though, in many cases in highly literate modern societies, writing itself can also be considered natural language.

Excluded from the definition of natural language are: artificial and constructed languages, such as those developed for works of fiction; languages of formal logic, such as those in computer programming; and non-human communication systems in nature, such as whale vocalizations or honey bees' waggle dance. The academic consensus is that particular key features prevent animal communication systems from being classified as languages at all. Certain human communication or linguistic systems with no native speakers, as sometimes used in cross-cultural contexts, are also not natural languages.

Waggle dance is a term used in beekeeping and ethology for a particular figure-eight dance of the honey bee. By performing this dance, successful foragers can share information about the direction and distance to patches of flowers yielding nectar and pollen, to water sources, or to new nest-site locations with other members of the colony.

The waggle dance and the round dance are two forms of dance behaviour that are part of a continuous transition. As the distance between the resource and the hive increases, the round dance transforms into variations of a transitional dance, which, when communicating resources at even greater distances, becomes the waggle dance. In the case of Apis mellifera ligustica, the round dance is performed until the resource is about ten metres (33 ft) away from the hive, transitional dances are performed when the resource is at a distance of twenty to thirty metres (66 to 98 ft) away from the hive, and finally, when it is located at distances greater than forty metres (130 ft) from the hive, the waggle dance is performed. However, even close to the nest, the round dance can contain elements of the waggle dance, such as a waggle portion. It has therefore been suggested that the term waggle dance is better for describing both the waggle dance and the round dance.

Mites are small arachnids (eight-legged arthropods) of two large orders, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari. However, most recent genetic analyses do not recover the two as each other's closest relative within Arachnida, rendering the group invalid as a clade. Most mites are tiny, less than 1 mm (0.04 in) in length, and have a simple, unsegmented body plan. The small size of most species makes them easily overlooked; some species live in water, many live in soil as decomposers, others live on plants, sometimes creating galls, while others are predators or parasites. This last type includes the commercially destructive Varroa parasite of honey bees, as well as scabies mites of humans. Most species are harmless to humans, but a few are associated with allergies or may transmit diseases.

The scientific discipline devoted to the study of mites is called acarology.

Ethology is a branch of zoology that studies the behaviour of non-human animals. It has its scientific roots in the work of Charles Darwin and of American and German ornithologists of the late 19th and early 20th century, including Charles O. Whitman, Oskar Heinroth, and Wallace Craig. The modern discipline of ethology is generally considered to have begun during the 1930s with the work of the Dutch biologist Nikolaas Tinbergen and the Austrian biologists Konrad Lorenz and Karl von Frisch, the three winners of the 1973 Nobel Prize in Physiology or Medicine. Ethology combines laboratory and field science, with a strong relation to neuroanatomy, ecology, and evolutionary biology.

Ecosystem services are the various benefits that humans derive from ecosystems. The interconnected living and non-living components of the natural environment offer benefits such as pollination of crops, clean air and water, decomposition of wastes, and flood control.

Ecosystem services are grouped into categories of services, which was popularized in the early 2000s by the Millennium Ecosystem Assessment (MA) initiative by the United Nations. How these groups are defined varies dependent on classification system. The MA groups the services into four broad categories of services. These are provisioning services, such as the production of food and water; regulating services, such as the control of climate and disease; supporting services, such as nutrient cycles and oxygen production; and cultural services, such as recreation, tourism, and spiritual gratification.

Bees are winged insects that form a monophyletic clade Anthophila within the superfamily Apoidea of the order Hymenoptera, with over 20,000 known species in seven recognized families. Some species – including honey bees, bumblebees, and stingless bees – are social insects living in highly hierarchical colonies, while over 90% of bee species – including mason bees, carpenter bees, leafcutter bees, and sweat bees – are solitary. Members of the most well-known bee genus, Apis (i.e. honey bees), are known to construct hexagonally celled waxy nests called hives.

Unlike the closely related wasps and ants, who are carnivorous/omnivorous, bees are herbivores that specifically feed on nectar (nectarivory) and pollen (palynivory), the former primarily as a carbohydrate source for metabolic energy, and the latter primarily for protein and other nutrients for their larvae. They are found on every continent except Antarctica, and in every habitat on the planet that contains insect-pollinated flowering plants. The most common bees in the Northern Hemisphere are the Halictidae, or sweat bees, but they are small and often mistaken for wasps or flies. Bees range in size from tiny stingless bee species, whose workers are less than 2 millimeters (0.08 in) long, to the leafcutter bee Megachile pluto, the largest species of bee, whose females can attain a length of 39 millimeters (1.54 in). Vertebrate predators of bees include primates and birds such as bee-eaters; insect predators include beewolves and dragonflies.

Apidae is the largest family within the superfamily Apoidea, containing at least 5700 species of bees. The family includes some of the most commonly seen bees, including bumblebees and honey bees, but also includes stingless bees (also used for honey production), carpenter bees, orchid bees, cuckoo bees, and a number of other less widely known groups. Many are valuable pollinators in natural habitats and for agricultural crops.

A beehive is an enclosed structure in which honey bees raise their young and produce honey as part of their seasonal cycle. Although the word beehive is used to describe the nest of any bee colony, scientific and professional literature distinguishes nest from hive. Nest is used to discuss colonies that house themselves in natural or artificial cavities or in structures that are hanging and exposed. The term hive is used to describe a man made structure created to house a honey bee colony. While species of Apis live in colonies, the western (Apis mellifera) and eastern honey bees (Apis cerana) are the main species kept in artificial beehives.

The hive's internal structure is a densely packed group of hexagonal prismatic cells made of beeswax, called a honeycomb. The bees use the cells to store food (honey and pollen) and to house the brood (eggs, larvae, and pupae).

A honeycomb is a mass of hexagonal prismatic cells built from beeswax by honey bees in their nests to contain their brood (eggs, larvae, and pupae) and stores of honey and pollen.

Beekeepers may remove the entire honeycomb to harvest honey. Honey bees consume about 8.4 lb (3.8 kg) of honey to secrete 1 lb (450 g) of wax, and so beekeepers may return the wax to the hive after harvesting the honey to improve honey outputs. The structure of the comb may be left basically intact when honey is extracted from it by uncapping and spinning in a centrifugal honey extractor. If the honeycomb is too worn out, the wax can be reused in a number of ways, including making sheets of comb foundation with a hexagonal pattern. Such foundation sheets allow the bees to build the comb with less effort, and the hexagonal pattern of worker-sized cell bases discourages the bees from building the larger drone cells. Fresh, new comb is sometimes sold and used intact as comb honey, especially if the honey is being spread on bread rather than used in cooking or as a sweetener.

Beeswax (also known as cera alba) is a natural wax produced by honey bees of the genus Apis. The wax is formed into scales by eight wax-producing glands in the abdominal segments of worker bees, which discard it in or at the hive. The hive workers collect and use it to form cells for honey storage and larval and pupal protection within the beehive. Chemically, beeswax consists mainly of esters of fatty acids and various long-chain alcohols.

Beeswax has been used since prehistory as the first plastic, as a lubricant and waterproofing agent, in lost wax casting of metals and glass, as a polish for wood and leather, for making candles, as an ingredient in cosmetics and as an artistic medium in encaustic painting.

Honey is a sweet and viscous substance made by several species of bees, the best-known of which are honey bees. Honey is made and stored to nourish bee colonies. Bees produce honey by gathering and then refining the sugary secretions of plants (primarily floral nectar) or the secretions of other insects, like the honeydew of aphids. This refinement takes place both within individual bees, through regurgitation and enzymatic activity, and during storage in the hive, through water evaporation that concentrates the honey's sugars until it is thick and viscous.

Honey bees stockpile honey in the hive. Within the hive is a structure made from wax called honeycomb. The honeycomb is made up of hundreds or thousands of hexagonal cells, into which the bees regurgitate honey for storage. Other honey-producing species of bee store the substance in different structures, such as the pots made of wax and resin used by the stingless bee.

The western honey bee or European honey bee (Apis mellifera) is the most common of the 7–12 species of honey bees worldwide. The genus name Apis is Latin for 'bee', and mellifera is the Latin for 'honey-bearing' or 'honey-carrying', referring to the species' production of honey.

Like all honey bee species, the western honey bee is eusocial, creating colonies with a single fertile female (or "queen"), many normally non-reproductive females or "workers", and a small proportion of fertile males or "drones". Individual colonies can house tens of thousands of bees. Colony activities are organized by complex communication between individuals, through both pheromones and the waggle dance.

Beekeeping (or apiculture, from Latin: apis + culture) is the maintenance of bee colonies, commonly in artificial beehives. Honey bees in the genus Apis are the most commonly kept species but other honey producing bees such as Melipona stingless bees are also kept. Beekeepers (or apiarists) keep bees to collect honey and other products of the hive: beeswax, propolis, bee pollen, and royal jelly. Other sources of beekeeping income include pollination of crops, raising queens, and production of package bees for sale. Bee hives are kept in an apiary or "bee yard".

The earliest evidence of humans collecting honey are from Spanish caves paintings dated 6,000 BCE, however it is not until 3,100 BCE that there is evidence from Egypt of beekeeping being practiced.

Apis cerana, the eastern honey bee, Asiatic honey bee or Asian honey bee, is a species of honey bee native to South, Southeast and East Asia. This species is the sister species of Apis koschevnikovi and both are in the same subgenus as the western (European) honey bee, Apis mellifera. A. cerana is known to live sympatrically along with Apis koschevnikovi within the same geographic location. Apis cerana colonies are known for building nests consisting of multiple combs in cavities containing a small entrance, presumably for defense against invasion by individuals of another nest. The diet of this honey bee species consists mostly of pollen and nectar, or honey. Moreover, Apis cerana is known for its highly social behavior, reflective of its classification as a type of honey bee.

The terms Apis cerana indica and Apis Indica or Indian honey bee, is an historic term, with all Asian hive bees now referred to as Apis cerana.

Varroa is a genus of parasitic mesostigmatan mites associated with honey bees, placed in its own family, Varroidae. The genus was named for Marcus Terentius Varro, a Roman scholar and beekeeper. The condition of a honeybee colony being infested with Varroa mites is called varroosis (also, incorrectly, varroatosis).

Varroa mites, specifically the species Varroa destructor, are recognised as the biggest pest to honeybees worldwide due to their ability to transmit diseases such as deformed wing virus (or DWV) to larval or pupating bees, resulting in death or severe deformity of the pupae.

Sexual maturity is the capability of an organism to reproduce.

Most multicellular organisms are unable to sexually reproduce at birth (animals) or germination (e.g. plants): depending on the species, it may be days, weeks, or years until they have developed enough to be able to do so; in addition, certain cues may trigger an organism to become sexually mature. These may be external, such as drought, or fire, that triggers sexual maturation of certain plants, or internal, such as percentage of body fat (certain animals). Internal cues are not to be confused with hormones, the chemical signals that initiate cellular processes leading to sexual maturity, but the production and secretion of hormones is triggered by such cues. In some species, immature males may delay sexual maturation in the presence of another sexually mature male, as in the male chicken (rooster), due to the intense, often lethal, combat engaged in by mature males. The female honeybee only becomes sexually mature if it is fed a special substance ("royal jelly") during the larval stage.

In beekeeping, bee brood or brood refers to the eggs, larvae and pupae of honeybees. The brood of Western honey bees develops within a bee hive. In man-made, removable frame hives, such as Langstroth hives, each frame which is mainly occupied by brood is called a brood frame. Brood frames usually have some pollen and nectar or honey in the upper corners of the frame. The rest of the brood frame cells may be empty or occupied by brood in various developmental stages. During the brood raising season, the bees may reuse the cells from which brood has emerged for additional brood or convert it to honey or pollen storage. Bees show remarkable flexibility in adapting cells to a use best suited for the hive's survival.