The Hominini (hominins) form a taxonomic tribe of the subfamily Homininae (hominines). They comprise two extant genera: Homo (humans) and Pan (chimpanzees and bonobos), and in standard usage exclude the genus Gorilla (gorillas), which is grouped separately within the subfamily Homininae.
The term Hominini was originally introduced by Camille Arambourg (1948), who combined the categories of Hominina and Simiina pursuant to Gray's classifications (1825).
Bipedalism is a form of terrestrial locomotion where an animal moves by means of its two rear (or lower) limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped /ˈbaɪpɛd/, meaning 'two feet' (from Latin bis 'double' and pes 'foot'). Types of bipedal movement include walking or running (a bipedal gait) and hopping.
Several groups of modern species are habitual bipeds whose normal method of locomotion is two-legged. In the Triassic period some groups of archosaurs, a group that includes crocodiles and dinosaurs, developed bipedalism; among the dinosaurs, all the early forms and many later groups were habitual or exclusive bipeds; the birds are members of a clade of exclusively bipedal dinosaurs, the theropods. Within mammals, habitual bipedalism has evolved multiple times, with the macropods, kangaroo rats and mice, springhare, hopping mice, pangolins and hominin apes such as australopithecines, including humans, as well as many other extinct groups evolving the trait independently.
Several expansions of populations of archaic humans (genus Homo) out of Africa and throughout Eurasia took place in the course of the Lower Paleolithic, and into the beginning Middle Paleolithic, between about 2.1 million and 0.2 million years ago (Ma).These expansions are collectively known as Out of Africa I, in contrast to the expansion of anatomically modern humans (Homo sapiens) into Eurasia, which may have begun shortly after 0.2 million years ago (known in this context as "Out of Africa II").
The earliest presence of Homo (or indeed any hominin) outside of Africa dates to close to 2 million years ago.A 2018 study identified possible hominin presence at Shangchen, central China, as early as 2.12 Ma based onmagnetostratigraphic dating of the lowest layer containing what may possibly be stone artefacts.The oldest known human skeletal remains outside of Africa are from Dmanisi, Georgia (Dmanisi skull 4), and are dated to 1.8 Ma. These remains are classified as Homo erectus georgicus.
The Paleolithic (/ˌpeɪlioʊˈlɪθɪk, ˌpæli-/ PAY-lee-oh-LITH-ik, PAL-ee-) or Old Stone Age is a period in human prehistory distinguished by the original development of stone tools. It represents almost the entire period of human prehistoric technology, extending from the earliest known use of stone tools by hominins, c. 3.3 million years ago, to the end of the Pleistocene, c. 11,650 cal BP.
The Paleolithic Age in Europe preceded the Mesolithic Age, although the date of the transition varies geographically by several thousand years. During the Paleolithic Age, hominins grouped together in small societies such as bands and subsisted by gathering plants, fishing, and hunting or scavenging wild animals. The Paleolithic Age is characterized by the use of knapped stone tools, although at the time humans also used wood and bone tools. Other organic commodities were adapted for use as tools, including leather and vegetable fibers; however, due to rapid decomposition, these have not survived to any great degree.
Behavioral modernity is a suite of behavioral and cognitive traits believed to distinguish current Homo sapiens from other anatomically modern humans, hominins, and primates. Most scholars agree that modern human behavior can be characterized by abstract thinking, planning depth, symbolic behavior (e.g., art, ornamentation), music and dance, exploitation of large game, and blade technologies, among others.
Underlying these behaviors and technological innovations are cognitive and cultural foundations that have been documented experimentally and ethnographically by evolutionary and cultural anthropologists. These human universal patterns include cumulative cultural adaptation, social norms, language, and extensive help and cooperation beyond close kin.
The Lower Paleolithic (or Lower Palaeolithic) is the earliest subdivision of the Paleolithic or Old Stone Age. It spans the time from around 3.3 million years ago when the first evidence for stone tool production and use by hominins appears in the current archaeological record, until around 300,000 years ago, spanning the Oldowan ("mode 1") and Acheulean ("mode 2") lithics industries.
In African archaeology, the time period roughly corresponds to the Early Stone Age, the earliest finds dating back to 3.3 million years ago, with Lomekwian stone tool technology, spanning Mode 1 stone tool technology, which begins roughly 2.6 million years ago and ends between 400,000 and 250,000 years ago, with Mode 2 technology.
Biological anthropology, also known as physical anthropology, is a natural science discipline concerned with the biological and behavioral aspects of human beings, their extinct hominin ancestors, and related non-human primates, particularly from an evolutionary perspective. This subfield of anthropology systematically studies human beings from a biological perspective.
The australopithecines (/ɒˈstrəloʊˈpɪθəsiːnz, ˈɔːstreɪloʊ-/), formally Australopithecina or Hominina, are generally any species in the related genera of Australopithecus and Paranthropus. It may also include members of Kenyanthropus, Ardipithecus, and Praeanthropus. The term comes from a former classification as members of a distinct subfamily, the Australopithecinae. They are classified within the Australopithecina subtribe of the Hominini tribe. These related species are sometimes collectively termed australopithecines, australopiths, or homininians. They are the extinct, close relatives of modern humans and, together with the extant genus Homo, comprise the human clade. There is no general agreement to whether australopithecines are closest relatives of modern humans, as it has been argued that they are more closely related to extant African apes. Members of the human clade, i.e. the Hominini after the split from the chimpanzees, are called Hominina (see Hominidae; terms "hominids" and hominins).
While none of the groups normally directly assigned to this group survived, the australopithecines do not appear to be literally extinct (in the sense of having no living descendants) as the genera Kenyanthropus, Paranthropus, and Homo probably emerged as sisters of a late Australopithecus species such as A. africanus and/or A. sediba.
Shangchen (Chinese: 上陈) is a Lower Palaeolithic archaeological site in Lantian County, Shaanxi, China, some 25 km south of Weinan.It was discovered in 1964, and excavated during 2004 and 2017.
Stone tools found at the site were dated based on magnetostratigraphy in a 2018 study. Possible artefacts were found in 17 layers, tentatively dated by their discoverers to between 1.26 Ma (palaeosol S15) and 2.12 Ma (loess L28). The date of 2.12 Ma predates the earliest known fossils of archaic humans in Eurasia (Homo erectus georgicus) by 300,000 years. Whether these potential tools were made by an early species in the genus Homo or another hominin species is unknown.
The Dmanisi hominins, Dmanisi people, or Dmanisi man were a population of Early Pleistocene hominins whose fossils have been recovered at Dmanisi, Georgia. The fossils and stone tools recovered at Dmanisi range in age from 1.85 to 1.77 million years old, making the Dmanisi hominins the earliest well-dated hominin fossils in Eurasia and the best preserved fossils of early Homo from a single site so early in time. Though their precise classification is controversial and disputed, the Dmanisi fossils are highly significant within research on early hominin migrations out of Africa. The Dmanisi hominins are known from over a hundred postcranial fossils and five famous well-preserved skulls, referred to as Dmanisi Skulls 1–5.
The taxonomic status of the Dmanisi hominins is somewhat unclear due to their small brain size, primitive skeletal architecture, and the range of variation exhibited between the skulls. Their initial description classified them as Homo (erectus?) ergaster (an otherwise African taxon), or potentially an early offshoot of later Asian H. erectus. The discovery of a massive jaw, D2600, in 2000 led researchers to hypothesize that more than one hominin taxon had been present at the site and in 2002, the jaw was designated as the type specimen of the new species Homo georgicus. Later analyses by the Dmanisi research team have concluded that all the skulls likely represent the same taxon with significant age-related and sexual dimorphism, though this is not a universally held view. In 2006, the team favoured subsuming the taxon under Homo erectus as H. erectus georgicus or H. e. ergaster georgicus. The nomenclature is still debated.