Evolutionary radiation in the context of "Echinoderm"

Marine life, sea life or ocean life is the collective ecological communities that encompass all aquatic animals, plants, algae, fungi, protists, single-celled microorganisms and associated viruses living in the saline water of marine habitats, either the sea water of marginal seas and oceans, or the brackish water of coastal wetlands, lagoons, estuaries and inland seas. As of 2023, more than 242,000 marine species have been documented, and perhaps two million marine species are yet to be documented. An average of 2,332 new species per year are being described. Marine life is studied scientifically in both marine biology and in biological oceanography.

By volume, oceans provide about 90% of the living space on Earth, and served as the cradle of life and vital biotic sanctuaries throughout Earth's geological history. The earliest known life forms evolved as anaerobic prokaryotes (archaea and bacteria) in the Archean oceans around the deep sea hydrothermal vents, before photoautotrophs appeared and allowed the microbial mats to expand into shallow water marine environments. The Great Oxygenation Event of the early Proterozoic significantly altered the marine chemistry, which likely caused a widespread anaerobe extinction event but also led to the evolution of eukaryotes through symbiogenesis between surviving anaerobes and aerobes. Complex life eventually arose out of marine eukaryotes during the Neoproterozoic, and which culminated in a large evolutionary radiation event of mostly sessile macrofaunae known as the Avalon Explosion. This was followed in the early Phanerozoic by a more prominent radiation event known as the Cambrian Explosion, where actively moving eumetazoan became prevalent. These marine life also expanded into fresh waters, where fungi and green algae that were washed ashore onto riparian areas started to take hold later during the Ordovician before rapidly expanding inland during the Silurian and Devonian, paving the way for terrestrial ecosystems to develop.

The Neoproterozoic is the last of the three geologic eras of the Proterozoic eon, spanning from 1 billion to 538.8 million years ago, and is the last era of the Precambrian "supereon". It is preceded by the Mesoproterozoic era and succeeded by the Paleozoic era of the Phanerozoic eon, and is further subdivided into three periods, the Tonian, Cryogenian and Ediacaran.

One of the most severe glaciation events known in the geologic record occurred during the Cryogenian period of the Neoproterozoic, when global ice sheets may have reached the equator and created a "Snowball Earth" lasting about 100 million years. The earliest fossils of complex life are found in the Tonian period in the form of Otavia, a primitive sponge, and the earliest fossil evidence of metazoan radiation are found in the Ediacaran period, which included the namesaked Ediacaran biota as well as the oldest definitive cnidarians and bilaterians in the fossil record.

The Ediacaran (/ˌiːdiˈækərən/ EE-dee-ACK-ər-ən; formerly Vendian) biota is a taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period (c. 635–538.8 Mya). These were enigmatic tubular and frond-shaped, mostly sessile, organisms. Trace fossils of these organisms have been found worldwide, and represent the earliest known complex multicellular organisms. The term "Ediacara biota" has received criticism from some scientists due to its alleged inconsistency, arbitrary exclusion of certain fossils, and inability to be precisely defined.

The Ediacaran biota may have undergone evolutionary radiation in a proposed event called the Avalon explosion, 575 million years ago. This was after the Earth had thawed from the Cryogenian period's extensive glaciation. This biota largely disappeared with the rapid increase in biodiversity known as the Cambrian explosion. Most of the currently existing body plans of animals first appeared in the fossil record of the Cambrian rather than the Ediacaran. For macroorganisms, the Cambrian biota appears to have almost completely replaced the organisms that dominated the Ediacaran fossil record, although relationships are still a matter of debate.

The Avalon explosion, named from the Precambrian faunal trace fossils discovered on the Avalon Peninsula in Newfoundland, eastern Canada, is a proposed evolutionary radiation of prehistoric animals about 575 million years ago in the Ediacaran period, with the Avalon explosion being one of three eras grouped in this time period. This evolutionary event is believed to have occurred some 33 million years earlier than the Cambrian explosion, which had been long thought to be when complex life started on Earth.

Scientists are still unsure of the full extent behind the development of the Avalon explosion, which resulted in a rapid increase in metazoan biodiversity, including the first appearance of some extant infrakingdoms/superphyla such as cnidarians and bilaterians. Many of the Avalon explosion animals are sessile soft-bodied organisms living in deep marine environments, and the first stages of the Avalon explosion were observed through comparatively minimal species.

The Phanerozoic is the current and the latest of the four geologic eons in the Earth's geologic time scale, covering the time period from 542 million years ago to the present. It is the eon during which abundant animal and plant life has proliferated, diversified and colonized various niches on the Earth's surface, beginning with the Cambrian period when animals first developed hard shells that can be clearly preserved in the fossil record. The time before the Phanerozoic, collectively called the Precambrian, is now divided into the Hadean, Archaean and Proterozoic eons.

The time span of the Phanerozoic starts with the sudden appearance of fossilised evidence of a number of animal phyla; the evolution of those phyla into diverse forms; the evolution of plants; the evolution of fish, arthropods and molluscs; the terrestrial colonization and evolution of insects, chelicerates, myriapods and tetrapods; and the development of modern flora dominated by vascular plants. During this time span, tectonic forces which move the continents had collected them into a single landmass known as Pangaea (the most recent supercontinent), which then separated into the current continental landmasses.

The Cambrian explosion (also known as Cambrian radiation or Cambrian diversification) is an interval of time beginning approximately 538.8 million years ago in the Cambrian period of the early Paleozoic, when a sudden radiation of complex life occurred and practically all major animal phyla started appearing in the fossil record. It lasted for about 13 to 25 million years and resulted in the divergence of most modern metazoan phyla. The event was accompanied by major diversification in other groups of organisms as well.

Before early Cambrian diversification, most organisms were relatively simple, composed of individual cells or small multicellular organisms, occasionally organized into colonies. As the rate of diversification subsequently accelerated, the variety of life became much more complex and began to resemble that of today. Almost all present-day animal phyla appeared during this period, including the earliest chordates.

The Silurian-Devonian Terrestrial Revolution, also known as the Devonian Plant Explosion (DePE) and the Devonian explosion, was a period of rapid colonization, diversification and radiation of land plants (particularly vascular plants) and fungi (especially dikaryans) on dry lands that occurred 428 to 359 million years ago (Mya) during the Silurian and Devonian periods, with the most critical phase occurring during the Late Silurian and Early Devonian.

This diversification of terrestrial photosynthetic florae had vast impacts on the biotic composition of the Earth's surface, especially upon the Earth's atmosphere by oxygenation and carbon fixation. Their roots also eroded into the rocks, creating a layer of water-holding and mineral/organic matter-rich soil on top of Earth's crust known as the pedosphere, and significantly altering the chemistry of Earth's lithosphere and hydrosphere. The floral activities following the Silurian-Devonian plant revolution also exerted significant influences on changes in the water cycle and global climate, as well as driving the biosphere by creating diverse layers of vegetations that provide both sustenance and refuge for both upland and wetland habitats, paving the way for all terrestrial and aquatic biomes that would follow.

The Devonian (/dəˈvoʊni.ən, dɛ-/ də-VOH-nee-ən, deh-) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.62 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.86 Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied.

The first significant evolutionary radiation of life on land occurred during the Devonian, as free-sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants (pteridospermatophytes) appeared. This rapid evolution and colonization process, which had begun during the Silurian, is known as the Silurian-Devonian Terrestrial Revolution. The earliest land animals, predominantly arthropods such as myriapods, arachnids and hexapods, also became well-established early in this period, after beginning their colonization of land at least from the Ordovician Period.

Cladistics (/kləˈdɪstɪks/ klə-DIST-iks; from Ancient Greek κλάδος kládos 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics (synapomorphies) that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms worms or fishes were used within a strict cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. Radiation results in the generation of new subclades by bifurcation, but in practice sexual hybridization may blur very closely related groupings.

As a hypothesis, a clade can be rejected only if some groupings were explicitly excluded. It may then be found that the excluded group did actually descend from the last common ancestor of the group, and thus emerged within the group. ("Evolved from" is misleading, because in cladistics all descendants stay in the ancestral group). To keep only valid clades, upon finding that the group is paraphyletic this way, either such excluded groups should be granted to the clade, or the group should be abolished.