Elasmobranch in the context of Most recent common ancestor


Elasmobranch in the context of Most recent common ancestor

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⭐ Core Definition: Elasmobranch

Elasmobranchii (/ɪˌlæzməˈbræŋki/) is a subclass of Chondrichthyes or cartilaginous fish, including modern sharks (division Selachii), and batomorphs (division Batomorphi, including rays, skates, and sawfish). Members of this subclass are characterised by having five to seven pairs of gill slits opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil.

The definition of the clade is unclear with respect to fossil chondrichthyans. Some authors consider it as equivalent to Neoselachii (the crown group clade including modern sharks, rays, and all other descendants of their last common ancestor). Other authors use the name Elasmobranchii for a broader branch-based group of all chondrichthyans more closely related to modern sharks and rays than to Holocephali (the clade containing chimaeras and their extinct relatives). Important extinct groups of elasmobranchs sensu lato include the hybodonts (Order Hybodontiformes), xenacanths (order Xenacanthformes) and Ctenacanthiformes. These are also often referred to as "sharks" in reference to their similar anatomy and ecology to modern sharks.

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Elasmobranch in the context of Shark

Sharks are a group of elasmobranch cartilaginous fishes characterized by a ribless endoskeleton, dermal denticles, five to seven gill slits on each side, and pectoral fins that are not fused to the head. Modern sharks are classified within the division Selachii and are the sister group to the Batomorphi (rays and skates). Some sources extend the term "shark" as an informal category including extinct members of Chondrichthyes (cartilaginous fish) with a shark-like morphology, such as hybodonts. Shark-like chondrichthyans such as Cladoselache and Doliodus first appeared in the Devonian Period (419–359 million years), though some fossilized chondrichthyan-like scales are as old as the Late Ordovician (458–444 million years ago). The earliest confirmed modern sharks (Selachii) are known from the Early Jurassic around 200 million years ago, with the oldest known member being Agaleus, though records of true sharks may extend back as far as the Permian.

Sharks range in size from the small dwarf lanternshark (Etmopterus perryi), a deep sea species that is only 17 centimetres (6.7 in) in length, to the whale shark (Rhincodon typus), the largest fish in the world, which reaches approximately 12 metres (40 ft) in length. They are found in all seas and are common to depths up to 2,000 metres (6,600 ft). They generally do not live in freshwater, although there are a few known exceptions, such as the bull shark and the river sharks, which can be found in both seawater and freshwater, and the Ganges shark, which lives only in freshwater. Sharks have a covering of placoid scales (denticles) that protects the skin from damage and parasites in addition to improving their fluid dynamics. They have numerous sets of replaceable teeth.

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Elasmobranch in the context of Dalatias licha

The kitefin shark or seal shark (Dalatias licha) is a species of squaliform shark in the family Dalatiidae, and the type species in its genus. It is found sporadically around the world, usually close to the sea floor at depths of 200–600 m (660–1,970 ft). With a sizable oil-filled liver to maintain neutral buoyancy, this shark is able to cruise slowly through the water while expending little energy. The kitefin shark, the largest luminous vertebrate on record, has a slender body with a very short, blunt snout, large eyes, and thick lips. Its teeth are highly differentiated between the upper and lower jaws, with the upper teeth small and narrow and the lower teeth large, triangular, and serrated. Its typical length is 1.0–1.4 m (3.3–4.6 ft), though examples as long as 5.9 ft (180 cm) have been encountered.

Armed with large teeth and a strong bite, the kitefin shark is a powerful, solitary predator that takes many different types of prey, ranging from bony fishes, sharks and rays, to cephalopods, crustaceans, polychaete worms, siphonophores, and possibly carrion. It also takes bites out of animals larger than itself, similar to its smaller relative, the cookiecutter shark (Isistius brasiliensis). This shark is aplacental viviparous and gives birth to 10–14 young. The kitefin shark is fished commercially for its meat, skin, and liver oil, primarily by Portugal and Japan. A fishery targeting this species existed off the Azores from the 1970s to the 1990s, but collapsed due to overfishing and falling liver oil prices; the rapid depletion of the Azores stock is often cited as an example of the susceptibility of deep-sea sharks to human exploitation. The low reproductive rate of this species renders it susceptible to overfishing and, coupled with known population declines, has led it to be assessed as Vulnerable by the International Union for Conservation of Nature (IUCN).

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