Diapsid in the context of Tuatara


Diapsid in the context of Tuatara

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⭐ Core Definition: Diapsid

Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The earliest traditionally identified diapsids, the araeoscelidians, appeared about three hundred million years ago during the late Carboniferous period. All diapsids other than the most primitive ones in the clade Araeoscelidia are often placed into the clade Neodiapsida. The diapsids are extremely diverse, and include birds and all modern reptile groups, including turtles, which were historically thought to lie outside the group. All modern reptiles and birds are placed within the neodiapsid subclade Sauria. Although some diapsids have lost either one hole (lizards), or both holes (snakes and turtles), or have a heavily restructured skull (modern birds), they are still classified as diapsids based on their ancestry. At least 17,084 species of diapsid animals are extant: 9,159 birds, and 7,925 snakes, lizards, tuatara, turtles, and crocodiles.

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Diapsid in the context of Archosaur

Archosauria (lit.'ruling reptiles') or archosaurs (/ˈɑːrkəˌsɔːr/) is a clade of diapsid sauropsid tetrapods, with birds and crocodilians being the only known extant representatives. Although broadly classified as reptiles, which traditionally exclude birds, the cladistic sense of the term includes all living and extinct relatives of birds and crocodilians such as non-avian dinosaurs, pterosaurs, phytosaurs, aetosaurs and rauisuchians as well as many Mesozoic marine reptiles. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants.

The base of Archosauria splits into two clades: Pseudosuchia, which includes crocodilians and their extinct relatives; and Avemetatarsalia, which includes birds and their extinct relatives (such as non-avian dinosaurs and pterosaurs). Older definitions of the group Archosauria rely on shared morphological characteristics, such as an antorbital fenestra in the skull, serrated teeth, and an upright stance. Some extinct reptiles, such as proterosuchids and euparkeriids, also possessed these features yet originated prior to the split between the crocodilian and bird lineages. The older morphological definition of Archosauria nowadays roughly corresponds to Archosauriformes, a group named to encompass crown-group archosaurs and their close relatives.

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Diapsid in the context of Avemetatarsalia

Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. Dinosaurs were the largest terrestrial animals for much of the Mesozoic Era, and one group of small feathered dinosaurs (Aves, i.e. birds) has survived up to the present day. Pterosaurs were the first flying vertebrates and persisted through the Mesozoic before dying out at the Cretaceous-Paleogene (K-Pg) extinction event. Both dinosaurs and pterosaurs appeared in the Triassic Period, shortly after avemetatarsalians as a whole. The name Avemetatarsalia was first established by British palaeontologist Michael Benton in 1999. An alternate name is Pan-Aves, or "all birds", in reference to its definition containing all animals, living or extinct, which are more closely related to birds than to crocodilians.

Although dinosaurs and pterosaurs were the only avemetatarsalians to survive past the end of the Triassic, other groups flourished during the Triassic. The most basal (earliest-branching) and plesiomorphic ("primitive") known avemetatarsalians were the aphanosaurs. Aphanosaurs were rare, four-legged carnivores which were only properly distinguished as a group in 2017. The split between dinosaurs and pterosaurs occurred just after aphanosaurs branched off the archosaur family tree. This split corresponds to the subgroup Ornithodira (Ancient Greek ὄρνις (órnis, "bird") + δειρή (deirḗ, "throat"), defined as the last common ancestor of dinosaurs and pterosaurs, and all of its descendants. Until the discovery of aphanosaurs, Ornithodira and Avemetatarsalia were considered roughly equivalent concepts.

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Diapsid in the context of Archosauriformes

Archosauriformes (Greek for 'ruling lizards', and Latin for 'form') is a clade of diapsid reptiles encompassing archosaurs and some of their close relatives. It was defined by Jacques Gauthier (1994) as the clade stemming from the last common ancestor of Proterosuchidae and Archosauria. Phil Senter (2005) defined it as the most exclusive clade containing Proterosuchus and Archosauria. Gauthier as part of the Phylonyms (2020) defined the clade as the last common ancestor of Gallus, Alligator, and Proterosuchus, and all its descendants. Archosauriforms are a branch of archosauromorphs which originated in the Late Permian (roughly 252 million years ago) and persist to the present day as the two surviving archosaur groups: crocodilians and birds.

Archosauriforms present several traits historically ascribed to the group Archosauria. These include serrated teeth set in deep sockets, a more active metabolism, and an antorbital fenestra (a hole in the skull in front of the eyes). Reptiles with these traits have also been termed "thecodonts" in older methods of classification. Thecodontia is a paraphyletic group, and its usage as a taxonomic category has been rejected under modern cladistic systems. The name Archosauriformes is intended as a monophyletic replacement compatible with modern taxonomy.

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Diapsid in the context of Temple (anatomy)

The temple, also known as the pterion, is a latch where four skull bones intersect: the frontal, parietal, temporal, and sphenoid. It is located on the side of the head behind the eye between the forehead and the ear. The temporal muscle covers this area and is used during mastication.

Cladistics classifies land vertebrates based on the presence of an upper hole, a lower hole, both, or neither in the cover of dermal bone that formerly covered the temporalis muscle, whose origin is the temple and whose insertion is the jaw.

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Diapsid in the context of Choristodera

Choristodera (from the Greek χωριστός chōristos + δέρη dérē, 'separated neck') is an extinct order of semiaquatic diapsid reptiles that ranged from the Middle Jurassic, or possibly Triassic, to the Miocene (168 to 20 or possibly 11.6 million years ago). Choristoderes are morphologically diverse, with the best known members being the crocodile-like neochoristoderes such as Champsosaurus. Other choristoderans had lizard-like or long necked morphologies. Choristoderes appear to have been confined to the Northern Hemisphere, having been found in North America, Asia, and Europe, and possibly also North Africa. Choristoderes are generally thought to be derived neodiapsids that are close relatives or members of Sauria.

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Diapsid in the context of Archosauromorpha

Archosauromorpha (Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) than to lepidosaurs (such as tuataras, lizards, and snakes). Archosauromorphs first appeared during the late Middle Permian or Late Permian, though they became much more common and diverse during the Triassic period.

Although Archosauromorpha was first named in 1946, its membership did not become well-established until the 1980s. Currently Archosauromorpha encompasses four main groups of reptiles: the stocky, herbivorous allokotosaurs and rhynchosaurs, the hugely diverse Archosauriformes, and a polyphyletic grouping of various long-necked reptiles including Protorosaurus, tanystropheids, and Prolacerta. Other groups including pantestudines (turtles and their extinct relatives) and the semiaquatic choristoderes have also been placed in Archosauromorpha by some authors.

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Diapsid in the context of Eu-

This is a list of common affixes used when scientifically naming species, particularly extinct species for whom only their scientific names are used, along with their derivations.

  • -ales: Pronunciation: /ˈa.lis/. Origin: Latin: -ālis. Meaning: Used to form taxonomic names of orders for plants and fungi.
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Diapsid in the context of Varanopidae

Varanopidae is an extinct family of amniotes known from the Late Carboniferous to Middle Permian that resembled monitor lizards (with the name of the group deriving from the monitor lizard genus Varanus) and may have filled a similar niche. Typically, they are considered to be relatively basal synapsids (and thus more closely related to mammals than to reptiles), although some studies from the late 2010s recovered them being taxonomically closer to diapsid reptiles, recent studies from the early 2020s support their traditional placement as synapsids on the basis of high degree of bone labyrinth ossification, maxillary canal morphology and phylogenetic analyses. A varanopid from the late Middle Permian Pristerognathus Assemblage Zone (Capitanian) is the youngest known varanopid and the last member of the "pelycosaur" group of synapsids. Thus, Varanopidae vanishes from the fossil record at the same time as dinocephalians, plausibly as a result of a major mass extinction event that has been called the "Dinocephalian extinction event".

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Diapsid in the context of Archosaurian

Archosauria or archosaurs (/ˈɑːrkəˌsɔːr/) is a clade of diapsid sauropsid tetrapods, with birds and crocodilians being the only known extant representatives. Although broadly classified as reptiles, which traditionally exclude birds, the cladistic sense of the term includes all living and extinct relatives of birds and crocodilians such as non-avian dinosaurs, pterosaurs, phytosaurs, aetosaurs and rauisuchians as well as many Mesozoic marine reptiles. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants.

The base of Archosauria splits into two clades: Pseudosuchia, which includes crocodilians and their extinct relatives; and Avemetatarsalia, which includes birds and their extinct relatives (such as non-avian dinosaurs and pterosaurs). Older definitions of the group Archosauria rely on shared morphological characteristics, such as an antorbital fenestra in the skull, serrated teeth, and an upright stance. Some extinct reptiles, such as proterosuchids and euparkeriids, also possessed these features yet originated prior to the split between the crocodilian and bird lineages. The older morphological definition of Archosauria nowadays roughly corresponds to Archosauriformes, a group named to encompass crown-group archosaurs and their close relatives.

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Diapsid in the context of Temporal fenestra

Temporal fenestrae are openings in the temporal region of the skull of some amniotes, behind the orbit (eye socket). These openings have historically been used to track the evolution and affinities of reptiles. Temporal fenestrae are commonly (although not universally) seen in the fossilized skulls of dinosaurs and other sauropsids (the total group of reptiles, including birds). The major reptile group Diapsida, for example, is defined by the presence of two temporal fenestrae on each side of the skull. The infratemporal fenestra, also called the lateral temporal fenestra or lower temporal fenestra, is the lower of the two and is exposed primarily in lateral (side) view.

The supratemporal fenestra, also called the upper temporal fenestra, is positioned above the other fenestra and is exposed primarily in dorsal (top) view. In some reptiles, particularly dinosaurs, the parts of the skull roof lying between the supratemporal fenestrae are thinned out by excavations from the adjacent fenestrae. These extended margins of thinned bone are called supratemporal fossae.

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Diapsid in the context of Araeoscelidia

Araeoscelidia or Araeoscelida is a clade of extinct tetrapods (traditionally classified as diapsid reptiles) superficially resembling lizards, extending from the Late Carboniferous to the Early Permian.The group contains the genera Araeoscelis, Petrolacosaurus, the possibly aquatic Spinoaequalis, and less well-known genera such as Kadaliosaurus and Zarcasaurus. This clade is usually considered to be the sister group to all (currently known) later diapsids.

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Diapsid in the context of Sauria

Sauria is the clade of diapsids containing the most recent common ancestor of Archosauria (which includes crocodilians and birds) and Lepidosauria (which includes squamates and the tuatara), and all its descendants. Since most molecular phylogenies recover turtles as more closely related to archosaurs than to lepidosaurs as part of Archelosauria, Sauria can be considered the crown group of diapsids, or reptiles in general. Depending on the systematics, Sauria includes all modern reptiles or most of them (including birds, a type of archosaur) as well as various extinct groups.

Sauria lies within the larger total group Sauropsida, which also contains various stem-reptiles which are more closely related to reptiles than to mammals. Prior to its modern usage, "Sauria" was used as a name for the suborder occupied by lizards, which before 1800 were considered crocodilians.

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Diapsid in the context of Euryapsid

Euryapsida is a polyphyletic (unnatural, as the various members are not closely related) group of sauropsids that are distinguished by a single temporal fenestra, an opening behind the orbit, under which the post-orbital and squamosal bones articulate. They are different from Synapsida, which also have a single opening behind the orbit, by the placement of the fenestra. In synapsids, this opening is below the articulation of the post-orbital and squamosal bones. It is now commonly believed that euryapsids (particularly sauropterygians) are in fact diapsids (which have two fenestrae behind the orbit) that lost the lower temporal fenestra. Euryapsids are usually considered entirely extinct, although turtles might be part of the sauropterygian clade while other authors disagree. Euryapsida may also be a synonym of Sauropterygia sensu lato.

The ichthyosaurian skull is sometimes described as having a metapsid (or parapsid) condition instead of a truly euryapsid one. In ichthyosaurs, the squamosal bone is never part of the fenestra's margin. Parapsida was originally a taxon consisting of ichthyosaurs, squamates, protorosaurs, araeoscelidans and pleurosaurs.

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Diapsid in the context of Anapsid

An anapsid is an amniote whose skull lacks one or more skull openings (fenestra, or fossae) near the temples. Traditionally, the Anapsida are considered the most primitive subclass of amniotes, the ancestral stock from which Synapsida and Diapsida evolved, making anapsids paraphyletic. It is, however, doubtful that all anapsids lack temporal fenestra as a primitive trait, and that all the groups traditionally seen as anapsids truly lacked fenestra.

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