The Zygnematales (Greek: ζυγός (zygós) and νῆμα (nḗma) (nom.), νήματος (nḗmatos) (gen.)), also called the Conjugatales, are an order of green algae, comprising several thousand different species in two families. The larger family Zygnemataceae, with well-known genera such as Zygnema and Spirogyra, includes members that grow as unbranched filaments, which grow longer through normal cell division. This group includes the desmids. Most members of both families live in freshwater, and form an important component of the algal scum that grows on or near plants and rocks.
Systematically they fall within the division Charophyta/Streptophyta, in which the land plants (Embryophyta) emerged.