Charales in the context of "Charophyceae"

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⭐ Core Definition: Charales

Charales is an order of freshwater green algae in the division Charophyta, class Charophyceae, commonly known as stoneworts. Depending on the treatment of the genus Nitellopsis, living (extant) species are placed into either one family (Characeae) or two (Characeae and Feistiellaceae). Further families are used for fossil members of the order. Linnaeus established the genus Chara in 1753.

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👉 Charales in the context of Charophyceae

Charophyceae is a class of charophyte green algae. AlgaeBase places it in division Charophyta. Extant (living) species are placed in a single order Charales, commonly known as "stoneworts" and "brittleworts". Fossil members of the class may be placed in separate orders, e.g. Sycidiales and Trochiliscales.

Charophyceae is basal in the Phragmoplastophyta clade which contains the embryophytes (land plants). In 2018, the first nuclear genome sequence from a species belonging to the Charophyceae was published: that of Chara braunii.

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Charales in the context of Charophyta

Charophyta (UK: /kəˈrɒfɪtə, ˌkærəˈftə/) is a paraphyletic group of freshwater green algae, called charophytes (/ˈkærəˌfts/), sometimes treated as a division, yet also as a superdivision. The terrestrial plants, the Embryophyta emerged deep within Charophyta, possibly from terrestrial unicellular charophytes, with the class Zygnematophyceae as a sister group.

With the Embryophyta now cladistically placed in the Charophyta, it is a synonym of Streptophyta. The sister group of the charophytes are the Chlorophyta. In some charophyte groups, such as the Zygnematophyceae or conjugating green algae, flagella are absent and sexual reproduction does not involve free-swimming flagellate sperm. Flagellate sperm, however, are found in stoneworts (Charales) and Coleochaetales, orders of parenchymatous charophytes that are the closest relatives of the land plants, where flagellate sperm are also present in all except the conifers and flowering plants. Fossil stoneworts of early Devonian age that are similar to those of the present day have been described from the Rhynie chert of Scotland. Somewhat different charophytes have also been collected from the Late Devonian (Famennian) Waterloo Farm lagerstätte of South Africa. These include two species each of Octochara and Hexachara, which are the oldest fossils of Charophyte axes bearing in situ oogonia.

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Charales in the context of Octochara

Octochara is a genus of fossil charophyte (aquatic green alga) from the Famennian (Late Devonian). It is one of two genera of charophyte described from the Waterloo Farm lagerstätte in southern Africa. It and Hexachara, from the same locality provide the oldest record of reconstruct able charophytes with in situ oogonia.

Octochara is derived from a Greek word "octo", meaning eight, a reference to the octoradial symmetry, and "chara", referring to membership of the Charales. In Octochara, a whorl of eight laterals are borne at each node. Each lateral is branched to produce four secondary branches and bears an oogonium.

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Charales in the context of Characeae

Characeae is a family of freshwater green algae in the order Charales, commonly known as stoneworts. They are also known as brittleworts or skunkweed, from the fragility of their lime-encrusted stems, and from the foul odor these produce when stepped on.

In some treatments, the Characeae includes all the living (extant) species of Charales; this circumscription is followed here. In other treatments, the genus Nitellopsis, which has both extant and extinct species, is placed in a separate family, Feistiellaceae.

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Charales in the context of Feistiellaceae

Feistiellaceae is a family of freshwater green algae in the order Charales.

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