Carbon fixation in the context of "Brown algae"

Archaea (/ɑːrˈkiːə/ ar-KEE-ə) is a domain of organisms. Traditionally, Archaea included only its prokaryotic members, but has since been found to be paraphyletic, as eukaryotes are known to have evolved from archaea. Even though the domain Archaea cladistically includes eukaryotes, the term archaea (sing. archaeon /ɑːrˈkiːɒn/ ar-KEE-on; from Ancient Greek ἀρχαῖον arkhaîon 'ancient') in English still generally refers specifically to prokaryotic members of Archaea. Archaea were initially classified as bacteria, receiving the name archaebacteria (/ˌɑːrkibækˈtɪəriə/, in the Archaebacteria kingdom), but this term has fallen out of use. Archaeal cells have unique properties separating them from Bacteria and Eukaryota, including: cell membranes made of ether-linked lipids; metabolisms such as methanogenesis; and a unique motility structure known as an archaellum. Archaea are further divided into multiple recognized phyla. Classification is difficult because most have not been isolated in a laboratory and have been detected only by their gene sequences in environmental samples. It is unknown if they can produce endospores.

Archaea are often similar to bacteria in size and shape, although a few have very different shapes, such as the flat, square cells of Haloquadratum walsbyi. Despite this, archaea possess genes and several metabolic pathways that are more closely related to those of eukaryotes, notably for the enzymes involved in transcription and translation. Other aspects of archaeal biochemistry are unique, such as their reliance on ether lipids in their cell membranes, including archaeols. Archaea use more diverse energy sources than eukaryotes, ranging from organic compounds such as sugars, to ammonia, metal ions or even hydrogen gas. The salt-tolerant Halobacteria use sunlight as an energy source, and other species of archaea fix carbon (autotrophy), but unlike cyanobacteria, no known species of archaea does both. Archaea reproduce asexually by binary fission, fragmentation, or budding; unlike bacteria, no known species of Archaea form endospores. The first observed archaea were extremophiles, living in extreme environments such as hot springs and salt lakes with no other organisms. Improved molecular detection tools led to the discovery of archaea in almost every habitat, including soil, oceans, and marshlands. Archaea are particularly numerous in the oceans, and the archaea in plankton may be one of the most abundant groups of organisms on the planet.

The Silurian-Devonian Terrestrial Revolution, also known as the Devonian Plant Explosion (DePE) and the Devonian explosion, was a period of rapid colonization, diversification and radiation of land plants (particularly vascular plants) and fungi (especially dikaryans) on dry lands that occurred 428 to 359 million years ago (Mya) during the Silurian and Devonian periods, with the most critical phase occurring during the Late Silurian and Early Devonian.

This diversification of terrestrial photosynthetic florae had vast impacts on the biotic composition of the Earth's surface, especially upon the Earth's atmosphere by oxygenation and carbon fixation. Their roots also eroded into the rocks, creating a layer of water-holding and mineral/organic matter-rich soil on top of Earth's crust known as the pedosphere, and significantly altering the chemistry of Earth's lithosphere and hydrosphere. The floral activities following the Silurian-Devonian plant revolution also exerted significant influences on changes in the water cycle and global climate, as well as driving the biosphere by creating diverse layers of vegetations that provide both sustenance and refuge for both upland and wetland habitats, paving the way for all terrestrial and aquatic biomes that would follow.

Photosynthesis (/ˌfoʊtəˈsɪnθəsɪs/ FOH-tə-SINTH-ə-sis) is a system of biological processes by which photopigment-bearing autotrophic organisms, such as most plants, algae and cyanobacteria, convert light energy — typically from sunlight — into the chemical energy necessary to fuel their metabolism. The term photosynthesis usually refers to oxygenic photosynthesis, a process that releases oxygen as a byproduct of water splitting. Photosynthetic organisms store the converted chemical energy within the bonds of intracellular organic compounds (complex compounds containing carbon), typically carbohydrates like sugars (mainly glucose, fructose and sucrose), starches, phytoglycogen and cellulose. When needing to use this stored energy, an organism's cells then metabolize the organic compounds through cellular respiration. Photosynthesis plays a critical role in producing and maintaining the oxygen content of the Earth's atmosphere, and it supplies most of the biological energy necessary for complex life on Earth.

Some organisms also perform anoxygenic photosynthesis, which does not produce oxygen. Some bacteria (e.g. purple bacteria) uses bacteriochlorophyll to split hydrogen sulfide as a reductant instead of water, releasing sulfur instead of oxygen, which was a dominant form of photosynthesis in the euxinic Canfield oceans during the Boring Billion. Archaea such as Halobacterium also perform a type of non-carbon-fixing anoxygenic photosynthesis, where the simpler photopigment retinal and its microbial rhodopsin derivatives are used to absorb green light and produce a proton (hydron) gradient across the cell membrane, and the subsequent ion movement powers transmembrane proton pumps to directly synthesize adenosine triphosphate (ATP), the "energy currency" of cells. Such archaeal photosynthesis might have been the earliest form of photosynthesis that evolved on Earth, as far back as the Paleoarchean, preceding that of cyanobacteria (see Purple Earth hypothesis).

Cyanobacteria (/saɪˌænoʊbækˈtɪəriə/ sy-AN-oh-bak-TEER-ee-ə) are a group of autotrophic gram-negative bacteria of the phylum Cyanobacteriota that can obtain biological energy via oxygenic photosynthesis. The name "cyanobacteria" (from Ancient Greek κύανος (kúanos) 'blue') refers to their bluish green (cyan) color, which forms the basis of cyanobacteria's informal common name, blue-green algae.

Cyanobacteria are probably the most numerous taxon to have ever existed on Earth and the first organisms known to have produced oxygen, having appeared in the middle Archean eon and apparently originated in a freshwater or terrestrial environment. Their photopigments can absorb the red- and blue-spectrum frequencies of sunlight (thus reflecting a greenish color) to split water molecules into hydrogen ions and oxygen. The hydrogen ions are used to react with carbon dioxide to produce complex organic compounds such as carbohydrates (a process known as carbon fixation), and the oxygen is released as a byproduct. By continuously producing and releasing oxygen over billions of years, cyanobacteria are thought to have converted the early Earth's anoxic, weakly reducing prebiotic atmosphere, into an oxidizing one with free gaseous oxygen (which previously would have been immediately removed by various surface reductants), resulting in the Great Oxidation Event and the "rusting of the Earth" during the early Proterozoic, dramatically changing the composition of life forms on Earth. The subsequent adaptation of early single-celled organisms to survive in oxygenous environments likely led to endosymbiosis between anaerobes and aerobes, and hence the evolution of eukaryotes during the Paleoproterozoic.

Zooplankton are the heterotrophic component of the planktonic community, having to consume other organisms to thrive. The name comes from Ancient Greek ζῷον (zōîon), meaning "animal", and πλαγκτός (planktós), meaning "drifter, wanderer, roamer", and thus, "animal drifter". Plankton are aquatic organisms that are unable to swim effectively against currents. Consequently, they drift or are carried along by currents in the ocean, or by currents in seas, lakes or rivers.

Zooplankton can be contrasted with phytoplankton (cyanobacteria and microalgae), which are the plant-like component of the plankton community (the "phyto-" prefix comes from Ancient Greek: φῠτόν, romanized: phutón, lit. 'plant', although taxonomically not plants). Zooplankton are heterotrophic (other-feeding), whereas phytoplankton are autotrophic (self-feeding), often generating biological energy and macromolecules through chlorophyllic carbon fixation using sunlight – in other words, zooplankton cannot manufacture their own food, while phytoplankton can. As a result, zooplankton must acquire nutrients by feeding on other organisms such as phytoplankton, which are generally smaller than zooplankton. Most zooplankton are microscopic but some (such as jellyfish) are macroscopic, meaning they can be seen with the naked eye.