A cambium (pl.: cambiums or cambia), in plants, is a tissue layer that provides partially undifferentiated cells for plant growth. It is found in the area between xylem and phloem. A cambium can also be defined as a cellular plant tissue from which phloem, xylem, or cork grows by division, resulting (in woody plants) in secondary thickening. It forms parallel rows of cells, which result in secondary tissues.
There are several distinct kinds of cambium found in plant stems and roots:
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Shield budding, also known as T-budding, is a technique of grafting to change varieties of fruit trees. Typically used in fruit tree propagation, it can also be used for many other kinds of nursery stock. An extremely sharp knife is necessary; specialty budding knives are on the market. A budding knife is a small knife with a type of spatula at the other end of the handle. The rootstock or stock plant may be cut off above the bud at budding, or one may wait until it is certain that the bud is growing.
Fruit tree budding is done when the bark "slips," i.e. the cambium is moist and actively growing. Rootstocks are young trees, either seedlings as Mazzard cherries for many cherry varieties, or clonal rootstocks (usually propagated by layering) when one wants highly consistent plants with well defined characteristics. The popular Malling-Merton series of rootstocks for apples was developed in England, and are used today for the majority of the commercial apple orchard trees.
In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips of stems and roots, causing them to elongate, and gives rise to primary tissue. Secondary growth occurs in most seed plants, but monocots usually lack secondary growth. If they do have secondary growth, it differs from the typical pattern of other seed plants.
The formation of secondary vascular tissues from the cambium is a characteristic feature of dicotyledons and gymnosperms. In certain monocots, the vascular tissues are also increased after the primary growth is completed but the cambium of these plants is of a different nature. In the living pteridophytes this feature is extremely rare, only occurring in Isoetes.
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