Breathing in the context of "Gill"

Exhalation (or expiration) is the flow of the breath out of an organism. In animals, it is the movement of air from the lungs out of the airways, to the external environment during breathing.This happens due to elastic properties of the lungs, as well as the internal intercostal muscles which lower the rib cage and decrease thoracic volume. As the thoracic diaphragm relaxes during exhalation it causes the tissue it has depressed to rise superiorly and put pressure on the lungs to expel the air. During forced exhalation, as when blowing out a candle, expiratory muscles including the abdominal muscles and internal intercostal muscles generate abdominal and thoracic pressure, which forces air out of the lungs.

Exhaled air is 4% carbon dioxide, a waste product of cellular respiration during the production of energy, which is stored as ATP. Exhalation has a complementary relationship to inhalation which together make up the respiratory cycle of a breath.

An aquatic animal is any animal, whether vertebrate or invertebrate, that lives in a body of water for all or most of its lifetime. Aquatic animals generally conduct aquatic respiration by extracting dissolved oxygen in water via specialised respiratory organs called gills, through the skin or across enteral mucosae, although some are secondarily aquatic animals (e.g. marine reptiles and marine mammals) evolved from terrestrial ancestors that re-adapted to aquatic environments, in which case they actually use lungs to breathe air and are essentially holding their breath when living in water. Some species of gastropod mollusc, such as the eastern emerald sea slug, are even capable of kleptoplastic photosynthesis via endosymbiosis with ingested yellow-green algae.

Almost all aquatic animals reproduce in water, either oviparously or viviparously, and many species routinely migrate between different water bodies during their life cycle. Some animals have fully aquatic life stages (typically as eggs and larvae), while as adults they become terrestrial or semi-aquatic after undergoing metamorphosis. Such examples include amphibians such as frogs, many flying insects such as mosquitoes, mayflies, dragonflies, damselflies and caddisflies, as well as some species of cephalopod molluscs such as the algae octopus (whose larvae are completely planktonic, but adults are highly terrestrial).

Hyperpnea, or hyperpnoea (forced respiration), is increased volume of air taken during breathing. It can occur with or without an increase in respiration rate. It is characterized by deep breathing. It may be physiologic—as when required by oxygen to meet metabolic demand of body tissues (for example, during or after heavy exercise, or when the body lacks oxygen at high altitude or as a result of anemia, or any other condition requiring more respiration)—or it may be pathologic, as when sepsis is severe or during pulmonary edema. Hyperpnea is further characterized by the required use of muscle contraction during both inspiration and expiration. Thus, hyperpnea is intense active breathing as opposed to the passive process of normal expiration.

Hyperpnea is distinguished from tachypnea, which is a respiratory rate greater than normal, resulting in rapid and shallow breaths, but not necessarily increasing volume in breathing. Hyperpnea is also distinguished from hyperventilation, which is over-ventilation (an increase in minute ventilation), which involves an increase in volume and respiration rate, resulting in rapid and deep breaths.

The respiratory system (also respiratory apparatus, ventilatory system) is a biological system consisting of specific organs and structures used for gas exchange in animals and plants.

In land animals, the respiratory surface is internalized as linings of the lungs. Gas exchange in the lungs occurs in millions of small air sacs. In mammals and reptiles, these are called alveoli, and in birds, they are known as atria. These microscopic air sacs have a rich blood supply, bringing the air into close contact with the blood. A system of airways, or hollow tubes, allow the air sacs to interface with the external environment; the largest of these is the trachea, which branches in the middle of the chest into the two main bronchi, which enter the lungs and branch into progressively narrower secondary and tertiary bronchi, which in turn branch into numerous smaller tubes known as the bronchioles in mammals and reptiles. In birds, the bronchioles are termed parabronchi. The bronchioles, or parabronchi, generally open into the microscopic alveoli (in mammals) and atria (in birds). Air has to be pumped from the environment into the alveoli or atria by the process of breathing which involves the muscles of respiration.

Pneuma (Ancient Greek: πνεῦμα, romanized: pneûma) is an ancient Greek word for "breath", and in a religious context for "spirit". It has various technical meanings for medical writers and philosophers of classical antiquity, particularly in regard to physiology, and is also used in Greek translations of ruach רוח in the Hebrew Bible, and in the Greek New Testament.

In classical philosophy, it is distinguishable from psyche (Ancient Greek: ψυχή, romanized: psūkhḗ), which originally meant "breath of life", but is regularly translated as "spirit" or most often "soul".

Fluid balance is an aspect of the homeostasis of organisms in which the amount of water in the organism needs to be controlled, via osmoregulation and behavior, such that the concentrations of electrolytes (salts in solution) in the various body fluids are kept within healthy ranges. The core principle of fluid balance is that the amount of water lost from the body must equal the amount of water taken in; for example, in humans, the output (via respiration, perspiration, urination, defecation, and expectoration) must equal the input (via eating and drinking, or by parenteral intake). Euvolemia is the state of normal body fluid volume, including blood volume, interstitial fluid volume, and intracellular fluid volume; hypovolemia and hypervolemia are imbalances. Water is necessary for all life on Earth. Humans can survive for 4 to 6 weeks without food but only for a few days without water.

Profuse sweating can increase the need for electrolyte replacement. Water-electrolyte imbalance produces headache and fatigue if mild; illness if moderate, and sometimes even death if severe. For example, water intoxication (which results in hyponatremia), the process of consuming too much water too quickly, can be fatal. Deficits to body water result in volume contraction and dehydration. Diarrhea is a threat to both body water volume and electrolyte levels, which is why diseases that cause diarrhea are great threats to fluid balance.

Several groups of tetrapods have undergone secondary aquatic adaptation, an evolutionary transition from being purely terrestrial to living at least partly aquatic. These animals are called "secondarily aquatic" because although all tetrapods descended from freshwater lobe finned fish (see evolution of tetrapods), their more recent ancestors are terrestrial vertebrates that evolved on land for hundreds of millions of years, and their clades only re-adapted to aquatic environment much later.

Unlike primarily aquatic vertebrates (i.e. fish), secondarily aquatic tetrapods (especially aquatic amniotes), while having appendages such as flippers, dorsal fin and tail fins (flukes) that resemble fish fins due to convergent evolution, still have physiology based on their terrestrial ancestry, most notably their air-breathing respiration via lungs (instead of aquatic respiration via gills) and excretion of nitrogenous waste as urea or uric acid (instead of ammonia like most fish). Nearly all extant aquatic tetrapods are secondarily aquatic, with only larval amphibians (tadpoles) being primarily aquatic with gills, and only some species of paedomorphic mole salamanders (most notably the fully aquatic axolotl) retain the gill-based physiology into adulthood.

Apnea (also spelled apnoea in British English) is the temporary cessation of breathing, which may be voluntary or involuntary. During apnea, there is no movement of the muscles of inhalation, and the volume of the lungs initially remains unchanged. Depending on how blocked the airways are (patency), there may or may not be a flow of gas between the lungs and the environment. If there is sufficient flow, gas exchange within the lungs and cellular respiration would not be severely affected. Voluntarily doing this is called "holding one's breath".Apnea may first be diagnosed in childhood, and it is recommended to consult an ear-nose-throat specialist, allergist or sleep physician to discuss symptoms when noticed; malformation and/or malfunctioning of the upper airways may be observed by an orthodontist.