Batoidea in the context of "Sawfish"

Sharks are a group of elasmobranch cartilaginous fishes characterized by a ribless endoskeleton, dermal denticles, five to seven gill slits on each side, and pectoral fins that are not fused to the head. Modern sharks are classified within the division Selachii and are the sister group to the Batomorphi (rays and skates). Some sources extend the term "shark" as an informal category including extinct members of Chondrichthyes (cartilaginous fish) with a shark-like morphology, such as hybodonts. Shark-like chondrichthyans such as Cladoselache and Doliodus first appeared in the Devonian Period (419–359 million years), though some fossilized chondrichthyan-like scales are as old as the Late Ordovician (458–444 million years ago). The earliest confirmed modern sharks (Selachii) are known from the Early Jurassic around 200 million years ago, with the oldest known member being Agaleus, though records of true sharks may extend back as far as the Permian.

Sharks range in size from the small dwarf lanternshark (Etmopterus perryi), a deep sea species that is only 17 centimetres (6.7 in) in length, to the whale shark (Rhincodon typus), the largest fish in the world, which reaches approximately 12 metres (40 ft) in length. They are found in all seas and are common to depths up to 2,000 metres (6,600 ft). They generally do not live in freshwater, although there are a few known exceptions, such as the bull shark and the river sharks, which can be found in both seawater and freshwater, and the Ganges shark, which lives only in freshwater. Sharks have a covering of placoid scales (denticles) that protects the skin from damage and parasites in addition to improving their fluid dynamics. They have numerous sets of replaceable teeth.

Cocos Island (Spanish: Isla del Coco) is a volcanic island in the Pacific Ocean administered by Costa Rica, approximately 550 km (342 mi; 297 nmi) southwest of the Costa Rican mainland. It constitutes the 11th of the 15 districts of Puntarenas Canton of the Province of Puntarenas. With an area of approximately 23.85 km (9.21 sq mi), the island is roughly rectangular in shape. It is the southernmost point of geopolitical North America if non-continental islands are included, and the only landmass above water on the Cocos tectonic plate.

The entirety of Cocos Island has been designated a Costa Rican National Park since 1978, and has no permanent inhabitants other than Costa Rican park rangers. While previously being portrayed as the largest uninhabited island within the tropics, this has been labeled a false claim, as the island of Fernandina in the Galapagos archipelago is also uninhabited and far larger in area. Surrounded by deep waters with counter-currents, Cocos Island is admired by scuba divers for its populations of hammerhead sharks, rays, dolphins and other large marine species. The wet climate and oceanic qualities give Cocos an ecological character that is not shared with either the Galápagos Archipelago or any of the other islands (for example, Malpelo, Gorgona or Coiba) in the eastern Pacific Ocean. Because of the unique ecology of the island and its surrounding waters, Cocos Island National Park became a UNESCO World Heritage Site in 1997. The island can only be reached by sea, which usually takes 36 to 48 hours.

Chondrichthyes (/kɒnˈdrɪkθiiːz/; from Ancient Greek χόνδρος (khóndros) 'cartilage' and ἰχθύς (ikhthús) 'fish') is a class of jawed fish that contains the cartilaginous fish or chondrichthyans, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or bony fish, which have skeletons primarily composed of bone tissue. Chondrichthyes are aquatic vertebrates with paired fins, paired nares, placoid scales, conus arteriosus in the heart, and a lack of opercula and swim bladders. Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates.

The class is divided into two subclasses: Elasmobranchii (sharks, rays, skates and sawfish) and Holocephali (chimaeras, sometimes called ghost sharks, which are sometimes separated into their own class). Extant chondrichthyans range in size from the 10 cm (3.9 in) finless sleeper ray to the over 10 m (33 ft) whale shark.

Acanthodii or acanthodians is an extinct class of gnathostomes (jawed fishes). They are currently considered to represent a paraphyletic grade of various fish lineages basal to extant Chondrichthyes, which includes living sharks, rays, and chimaeras. Acanthodians possess a mosaic of features shared with both osteichthyans (bony fish) and chondrichthyans (cartilaginous fish). In general body shape, they were similar to modern sharks, but their epidermis was covered with tiny rhomboid platelets like the scales of holosteians (gars, bowfins).

The popular name "spiny sharks" is because they were superficially shark-shaped, with a streamlined body, paired fins, a strongly upturned tail, and stout, largely immovable bony spines supporting all the fins except the tail—hence, "spiny sharks". However, acanthodians are not true sharks; their close relation to modern cartilaginous fish can lead them to be considered "stem-sharks". Acanthodians had a cartilaginous skeleton, but their fins had a wide, bony base and were reinforced on their anterior margin with a dentine spine. As a result, fossilized spines and scales are often all that remains of these fishes in ancient sedimentary rocks. The earliest acanthodians were marine, but during the Devonian, freshwater species became predominant.

Elasmobranchii (/ɪˌlæzməˈbræŋkiaɪ/) is a subclass of Chondrichthyes or cartilaginous fish, including modern sharks (division Selachii), and batomorphs (division Batomorphi, including rays, skates, and sawfish). Members of this subclass are characterised by having five to seven pairs of gill slits opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil.

The definition of the clade is unclear with respect to fossil chondrichthyans. Some authors consider it as equivalent to Neoselachii (the crown group clade including modern sharks, rays, and all other descendants of their last common ancestor). Other authors use the name Elasmobranchii for a broader branch-based group of all chondrichthyans more closely related to modern sharks and rays than to Holocephali (the clade containing chimaeras and their extinct relatives). Important extinct groups of elasmobranchs sensu lato include the hybodonts (Order Hybodontiformes), xenacanths (order Xenacanthformes) and Ctenacanthiformes. These are also often referred to as "sharks" in reference to their similar anatomy and ecology to modern sharks.

A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as well as possible hydrodynamic advantages. The term scale derives from the Old French escale, meaning a shell pod or husk.

Scales vary enormously in size, shape, structure, and extent, ranging from strong and rigid armour plates in fishes such as shrimpfishes and boxfishes, to microscopic or absent in fishes such as eels and anglerfishes. The morphology of a scale can be used to identify the species of fish it came from. Scales originated within the jawless ostracoderms, ancestors to all jawed fishes today.Most bony fishes are covered with the cycloid scales of salmon and carp, or the ctenoid scales of perch, or the ganoid scales of sturgeons and gars. Cartilaginous fishes (sharks and rays) are covered with placoid scales. Some species are covered instead by scutes, and others have no outer covering on part or all of the skin.

Gill slits are individual openings to gills, i.e., multiple gill arches, which lack a single outer cover. Such gills are characteristic of cartilaginous fish such as sharks and rays, as well as deep-branching vertebrates such as lampreys. In contrast, bony fishes have a single outer bony gill covering called an operculum.

Most sharks and rays have five pairs of gill slits, but a few species have 6 or 7 pairs. Shark gill slits lie in a row behind the head. The anterior edge of a gill slit is motile, moving outward to allow water to exit, but closing to prevent reverse flow. A modified slit, called a spiracle, lies just behind the eye, which assists the shark with taking in water during respiration and plays a major role in bottom–dwelling sharks. Spiracles are reduced or missing in active pelagic sharks. While the shark is moving, water passes through the mouth and over the gills in a process known as "ram ventilation". While at rest, most sharks pump water over their gills to ensure a constant supply of oxygenated water. A small number of species have lost the ability to pump water through their gills and must swim without rest. These species are obligate ram ventilators and would presumably asphyxiate if unable to move. Obligate ram ventilation is also true of some pelagic bony fish species.

Skates are cartilaginous fish belonging to the family Rajidae in the superorder Batoidea of rays. More than 150 species have been described, in 17 genera. Softnose skates and pygmy skates were previously treated as subfamilies of Rajidae (Arhynchobatinae and Gurgesiellinae), but are now considered as distinct families. Alternatively, the name "skate" is used to refer to the entire order of Rajiformes (families Anacanthobatidae, Arhynchobatidae, Gurgesiellidae and Rajidae).

Members of Rajidae are distinguished by a stiff snout and a rostrum that is not reduced.