Aposematism in the context of "Müllerian mimic"

In evolutionary biology, mimicry is the evolved resemblance of an organism to something else, often another organism of a different species. Mimicry may evolve between different species, or between individuals of the same species. In the simplest case, as in Batesian mimicry, a mimic resembles a model, so as to deceive a dupe, all three being of different species. A Batesian mimic, such as a hoverfly, is harmless, while its model, such as a wasp, is harmful, and is avoided by the dupe, such as an insect-eating bird. Birds hunt by sight, so the mimicry in that case is visual, but in other cases mimicry may make use of any of the senses. Most types of mimicry, including Batesian, are deceptive, as the mimics are not harmful, but Müllerian mimicry, where different harmful species resemble each other, is honest, as when species of wasps and of bees all have genuinely aposematic warning coloration. More complex types may be bipolar, involving only two species, such as when the model and the dupe are the same; this occurs for example in aggressive mimicry, where a predator in wolf-in-sheep's-clothing style resembles its prey, allowing it to hunt undetected. Mimicry is not limited to animals; in Pouyannian mimicry, an orchid flower is the mimic, resembling a female bee, its model; the dupe is the male bee of the same species, which tries to copulate with the flower, enabling it to transfer pollen, so the mimicry is again bipolar. In automimicry, another bipolar system, model and mimic are the same, as when blue lycaenid butterflies have 'tails' or eyespots on their wings that mimic their own heads, misdirecting predator dupes to strike harmlessly. Many other types of mimicry exist.

Patterns in nature are visible regularities of form found in the natural world. These patterns recur in different contexts and can sometimes be modelled mathematically. Natural patterns include symmetries, trees, spirals, meanders, waves, foams, tessellations, cracks and stripes. Early Greek philosophers studied pattern, with Plato, Pythagoras and Empedocles attempting to explain order in nature. The modern understanding of visible patterns developed gradually over time.

In the 19th century, the Belgian physicist Joseph Plateau examined soap films, leading him to formulate the concept of a minimal surface. The German biologist and artist Ernst Haeckel painted hundreds of marine organisms to emphasise their symmetry. Scottish biologist D'Arcy Thompson pioneered the study of growth patterns in both plants and animals, showing that simple equations could explain spiral growth. In the 20th century, the British mathematician Alan Turing predicted mechanisms of morphogenesis which give rise to patterns of spots and stripes. The Hungarian biologist Aristid Lindenmayer and the French American mathematician Benoît Mandelbrot showed how the mathematics of fractals could create plant growth patterns.

Skunks are mammals in the family Mephitidae. They are known for their ability to spray a liquid with a strong, unpleasant scent from their anal glands. Different species of skunk vary in appearance from black-and-white to brown, cream or ginger colored, but all have warning coloration.

While related to polecats and other members of the weasel family, skunks have as their closest relatives the Old World stink badgers.

Aggression is behavior aimed at opposing or attacking something or someone. Though often done with the intent to cause harm, some might channel it into creative and practical outlets. It may occur either reactively or without provocation. In humans, aggression can be caused by various triggers. For example, built-up frustration due to blocked goals or perceived disrespect. Human aggression can be classified into direct and indirect aggression; while the former is characterized by physical or verbal behavior intended to cause harm to someone, the latter is characterized by behavior intended to harm the social relations of an individual or group.

In definitions commonly used in the social sciences and behavioral sciences, aggression is an action or response by an individual that delivers something unpleasant to another person. Some definitions include that the individual must intend to harm another person.

Batesian mimicry is a form of mimicry wherein a harmless species has evolved to imitate the warning signals of a harmful species in order to benefit from these signals' tendency to deter their mutual predators. It is named after the English naturalist Henry Walter Bates, who worked on butterflies in the rainforests of Brazil.

Batesian mimicry is the most commonly-known and widely-studied of the mimicry complexes, such that the word "mimicry" is often synonymous with the type described by Batesian mimicry. However, there are many other forms of mimicry, some very similar in principle to the Batesian variety, others far separated.

Müllerian mimicry is a type of biological mimicry in which two or more well-defended species, often foul-tasting and sharing common predators, converge in appearance to mimic each other's honest warning signals. This convergence of appearance achieves the following benefit to species that undergo it: predators need only experience a single unpleasant encounter with any member of a set of Müllerian mimics in order to thereafter avoid all creatures of similar appearance, whether or not it is the same species as the initial encounter. A ring of distinct species is thereby protected from their mutual predators by attempted predation upon any one of its members. The phenomenon is named after the German-Brazilian naturalist Fritz Müller, who proposed the concept in 1878, supporting his theory with a mathematical model of frequency-dependent selection, one of the first such models anywhere in biology.

Müllerian mimicry was first identified in tropical butterflies that shared colourful wing patterns, but it is found in many groups of insects such as bumblebees, as well as in other animals such as poison frogs and coral snakes. The mimicry need not be restricted to that detected by vision—many snakes share auditory warning signals. Similarly, the defences involved are not limited to toxicity—anything that tends to deter predators, such as foul taste, sharp spines, or defensive behaviour can make a species unprofitable enough to predators to allow Müllerian mimicry to develop.

In zoology, automimicry, Browerian mimicry, or intraspecific mimicry, is a form of mimicry in which the same species of animal is imitated. There are two different forms.

In one form, first described by Lincoln Brower in 1967, weakly-defended members of a species with warning coloration are parasitic on more strongly-defended members of their species, mimicking them to provide the negative reinforcement learning required for warning signals to function. The mechanism, analogous to Batesian mimicry, is found in insects such as the monarch butterfly.

Animal coloration is the general appearance of an animal resulting from the reflection or emission of light from its surfaces. Some animals are brightly coloured, while others are hard to see. In some species, such as the peafowl, the male has strong patterns, conspicuous colours and is iridescent, while the female is far less visible.

There are several separate reasons why animals have evolved colours. Camouflage enables an animal to remain hidden from view. Animals use colour to advertise services such as cleaning to animals of other species; to signal their sexual status to other members of the same species; and in mimicry, taking advantage of the warning coloration of another species. Some animals use flashes of colour to divert attacks by startling predators. Zebras may possibly use motion dazzle, confusing a predator's attack by moving a bold pattern rapidly. Some animals are coloured for physical protection, with pigments in the skin to protect against sunburn, while some frogs can lighten or darken their skin for temperature regulation. Finally, animals can be coloured incidentally. For example, blood is red because the haem pigment needed to carry oxygen is red. Animals coloured in these ways can have striking natural patterns.