Ammonoidea in the context of "Catanduanes"

A cephalopod /ˈsɛfələpɒd/ is any member of the molluscan class Cephalopoda /sɛfəˈlɒpədə/ (Greek plural κεφαλόποδες, kephalópodes; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers and Swedes sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology.

Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by Nautilus and Allonautilus. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been identified. Two important extinct taxa are the Ammonoidea (ammonites) and Belemnoidea (belemnites). Extant cephalopods range in size from the 10 mm (0.3 in) Idiosepius thailandicus to the 700 kilograms (1,500 lb) heavy colossal squid, the largest extant invertebrate.

Nautiloids are a group of cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Fossil nautiloids are diverse and species rich, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). No orthoconic and only a handful of coiled species, the nautiluses, survive to the present day.

In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea sensu lato). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant relationships, this is not the case for nautiloids. Instead, nautiloids are a paraphyletic grade of various early-diverging cephalopod lineages, including the ancestors of ammonoids and coleoids. Some authors prefer a narrower definition of Nautiloidea (Nautiloidea sensu stricto), as a singular subclass including only those cephalopods which are closer to living nautiluses than they are to either ammonoids or coleoids.

Coleoidea or Dibranchiata is one of the two subclasses of cephalopod molluscs containing all the various taxa popularly thought of as "soft-bodied" or "shell-less" (i.e. octopus, squid and cuttlefish). Unlike its sister groups, the shelled †Ammonoidea and Nautiloidea, the coleoids have at most an internal shell called cuttlebone or gladius that is used for buoyancy or as muscle attachment. Some species, notably the incirrate octopuses (including commonly known varieties living in the shallows), have lost their internal shell altogether, while in some it has been replaced by a chitinous support structure.

The Early Devonian is the first of three epochs comprising the Devonian period, corresponding to the Lower Devonian series. It lasted from 419.62 ± 1.36 to 393.47 ± 0.99 and began with the Lochkovian Stage 419.62 ± 1.36 to 413.02 ± 0.99, which was followed by the Pragian from 413.02 ± 1.36 to 410.62 ± 0.99 and then by the Emsian, which lasted until the Middle Devonian began, 393.47± 0.99 million years ago.During this time, the first ammonoids appeared, descending from bactritoid nautiloids. Ammonoids during this time period were simple and differed little from their nautiloid counterparts. These ammonoids belong to the order Agoniatitida, which in later epochs evolved to new ammonoid orders, for example Goniatitida and Clymeniida. This class of cephalopod molluscs would dominate the marine fauna until the beginning of the Mesozoic Era.

The Carnian pluvial episode (CPE), often called the Carnian pluvial event, was a period of major change in global climate that coincided with significant changes in Earth's biota both in the sea and on land. It occurred during the latter part of the Carnian Stage, the first subdivision of the Late Triassic Epoch, and lasted for perhaps 1–2 million years (around 234–232 million years ago). Volcanic activity off the coast of North America led to global warming and increased rainfall on land, alongside a reduction of carbonate platforms in the oceans. Pluvial means "of or relating to rain; characterized by much rain, rainy."

The CPE corresponds to a significant episode in the evolution and diversification of many taxa that are important today. The earliest dinosaurs (which include the ancestors of birds), lepidosaurs (the ancestors of modern-day lizards, snakes, and the tuatara) and potentially mammaliaforms (ancestors of mammals) all diversified during the event. In the marine realm it saw the first appearance among the microplankton of coccoliths and dinoflagellates, with the latter linked to the rapid diversification of scleractinian corals through the establishment of symbiotic zooxanthellae within them. The CPE also saw the extinction of many aquatic invertebrate species, especially among ammonoids, bryozoans, and crinoids.