Western Steppe Herder in the context of Eastern Hunter-Gatherer


Western Steppe Herder in the context of Eastern Hunter-Gatherer

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⭐ Core Definition: Western Steppe Herder

In archaeogenetics, the term Western Steppe Herders (WSH), or Western Steppe Pastoralists, is the name given to a distinct ancestral component first identified in individuals from the Chalcolithic steppe around the start of the 5th millennium BC, subsequently detected in several genetically similar or directly related ancient populations including the Khvalynsk, Repin, Sredny Stog, and Yamnaya cultures, and found in substantial levels in contemporary populations of Europe, Central Asia, West Asia, and South Asia. This ancestry is often referred to as Yamnaya ancestry, Yamnaya-related ancestry, Steppe ancestry, or Steppe-related ancestry.

Western Steppe Herders are considered to be descended from a merger between Eastern Hunter-Gatherers (EHGs) and Caucasus Hunter-Gatherers (CHGs). The WSH component is modeled as an admixture of EHG and CHG ancestral components in roughly equal proportions, with the majority of the Y-DNA haplogroup contribution from EHG males. The Y-DNA haplogroups of Western Steppe Herder males are not uniform, with the Yamnaya culture individuals mainly belonging to R1b-Z2103 with a minority of I2a2, the earlier Khvalynsk culture also with mainly R1b but also some R1a, Q1a, J, and I2a2, and the later, high WSH ancestry Corded Ware culture individuals mainly belonging to haplogroup R1b in the earliest samples, with R1a-M417 becoming predominant over time.

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Western Steppe Herder in the context of Caucasus Hunter-Gatherer

Caucasus hunter-gatherer (CHG), also called Satsurblia cluster, is an anatomically modern human genetic lineage, first identified in a 2015 study, based on the population genetics of several modern Western Eurasian (European, Caucasian and Near Eastern) populations.

It represents an ancestry maximised in some Upper Paleolithic and Mesolithic hunter-gatherer groups in the Caucasus. These groups are also very closely related to Mesolithic hunter-gatherers and Neolithic farmers and pastoralists in the Iranian Plateau (Iranian hunter-gatherer cluster), who are sometimes included within the CHG group. Ancestry that is closely related to CHG-Iranian hunter gatherers and farmers is also known from further east, including from the Bactria–Margiana Archaeological Complex and the Harappan/Indus Valley Civilisation. Caucasus hunter-gatherers and Eastern hunter-gatherers are ancestral in roughly equal proportions to the Western Steppe Herders (WSH), who were widely spread across Europe and Asia beginning during the Chalcolithic.

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Western Steppe Herder in the context of Tarim mummies

The Tarim mummies are a series of mummies discovered in the Tarim Basin in present-day Xinjiang, China, which date from 1800 BCE to the first centuries BCE, with a new group of individuals recently dated to between c. 2100 and 1700 BCE. The Tarim population to which the earliest mummies belonged was agropastoral, and they lived c. 2000 BCE in what was formerly a freshwater environment, which has now become desertified.

Zhang et al. (2021) found that these early mummies (dating from 2,135 to 1,623 BCE) had high levels of Ancient North Eurasian ancestry (ANE, about 72%), with smaller admixture from Ancient Northeast Asians (ANA, about 28%), but no detectable Western Steppe-related ancestry. They formed a genetically isolated local population that "adopted neighbouring pastoralist and agriculturalist practices, which allowed them to settle and thrive along the shifting riverine oases of the Taklamakan Desert." These mummified individuals were long suspected to have been "Proto-Tocharian-speaking pastoralists", related to the Afanasievo or BMAC cultures, but "the earliest Tarim Basin cultures appear to have arisen from a genetically isolated local population." Zhang et al. (2025) investigated a Late Bronze Age site in the far west of the Tarim Basin, dated 1600 to 1400 BC. Its inhabitants overwhelmingly descended from the Sintashta and Andronovo population, with additional ancestry from BMAC (10%) and Tarim_EMBA (12%). Nearly all subjects belonged to Y-DNA haplogroup R-M17.

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