Sparassodonta in the context of Florentino Ameghino


Sparassodonta in the context of Florentino Ameghino

⭐ Core Definition: Sparassodonta

Sparassodonta (from Greek σπαράσσειν [sparassein], to tear, rend; and ὀδούς, gen. ὀδόντος [odous, odontos], tooth) is an extinct order of carnivorous metatherian mammals native to South America, related to modern marsupials. They were once considered to be true marsupials, but are now thought to be a separate side branch that split before the last common ancestor of all modern marsupials.

A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless terror birds. Previously, it was thought that these mammals died out in the face of competition from "more competitive" placental carnivorans during the Pliocene Great American Interchange, but more recent research has shown that sparassodonts died out long before eutherian carnivores arrived in South America (aside from procyonids, which sparassodonts probably did not directly compete with).

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Sparassodonta in the context of Machairodontinae

Machairodontinae (from Ancient Greek μάχαιρα (mákhaira), a type of ancient sword, and ὀδούς (odoús), meaning "tooth") is an extinct subfamily of carnivoran mammals of the cat family Felidae, representing the earliest diverging major branch of the family.

Machairodonts varied in size from comparable to lynxes to exceeding that of lions. The Machairodontinae contain many of the extinct predators commonly known as "saber-toothed cats", including those with greatly elongated upper maxillary canines, such as the famed genus Smilodon and Megantereon, though the degree of elongation was variable, and in some machairodontines like Dinofelis the length of the upper canines was much more modest. Sometimes, other carnivorous mammals with elongated teeth are also called saber-toothed cats, although they do not belong to the felids. Besides the machairodonts, other saber-toothed predators also arose in the nimravids, barbourofelids, machaeroidines, hyaenodonts and even in two groups of metatherians (the thylacosmilid sparassodonts and the deltatheroideans). Unlike living big cats, which generally clamp the muzzle or throat of prey to asphyxiate them, saber-toothed machairodontines are thought to have killed prey using a bite to the neck once immobilised, using their neck muscles to drive the saber teeth into the throat while the lower jaw served as an anchor, causing rapid death via blood loss.

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Sparassodonta in the context of Saber-toothed predator

A saber-tooth (alternatively spelled sabre-tooth) is any member of various extinct groups of predatory therapsids, predominantly carnivoran mammals, that are characterized by long, curved saber-shaped canine teeth which protruded from the mouth when closed.

Among the earliest animals that can be described as "sabertooths" are the gorgonopsids, a group of non-mammalian therapsids that lived during the Middle-Late Permian, around 270-252 million years ago. Saber-toothed mammals have been found almost worldwide from the Eocene epoch to the end of the Pleistocene epoch (42 million years ago – 11,000 years ago).

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Sparassodonta in the context of Thylacosmilidae

Thylacosmilidae is an extinct family of metatherian predators, related to the modern marsupials, which lived in South America between the Miocene and Pliocene epochs. Like other South American mammalian predators that lived prior to the Great American Biotic Interchange, these animals belonged to the order Sparassodonta, which occupied the ecological niche of many eutherian mammals of the order Carnivora from other continents. The family's most notable feature are the elongated, laterally flattened fangs, which is a remarkable evolutionary convergence with other saber-toothed mammals like Barbourofelis and Smilodon.

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Sparassodonta in the context of Deltatheroida

Deltatheroida is an extinct group of basal metatherians that were distantly related to modern marsupials. The majority of known members of the group lived in the Cretaceous; one species, Gurbanodelta kara, is known from the late Paleocene (Gashatan) of China. Their fossils are restricted to Central Asia and North America. This order can be defined as all metatherians closer to Deltatheridium than to Marsupialia.

When they were first identified in the 1920s, they were believed to be placentals and possible ancestors of the "creodonts" (a polyphyletic group of extinct carnivorous mammals from the Paleogene and Miocene), but this was later disproven. Nonetheless, deltatheroideans do converge on hyaenodontids, oxyaenids, carnivorans, dasyuromorphs, thylacoleonids and sparassodonts in many details of their dental anatomy, suggesting a carnivorous lifestyle.

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