Social insects in the context of Reproductive

The concept of the struggle for existence (or struggle for life) concerns the competition or battle for resources needed to live. It can refer to human society, or to organisms in nature. The concept is ancient, and the term struggle for existence was in use by the end of the 18th century. From the 17th century onwards the concept was associated with a population exceeding resources, an issue shown starkly in Thomas Robert Malthus’ An Essay on the Principle of Population which drew on Benjamin Franklin's Observations Concerning the Increase of Mankind, Peopling of Countries, etc..

It is sometimes forgotten that Charles Darwin used the term "struggle for existence" in what he called a metaphorical sense. This was because he used it to refer not only to direct competition between (and within) species but indirect competition (as with a plant at the edge of the desert). Darwin noted that the struggle for existence could also involve active or passive mutual aid between organisms of the same or different species, instancing social insects (see also symbiosis).

A nest is a structure built by certain animals to hold their eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic material such as twigs, grass, and leaves, or may be a simple depression in the ground, or a hole in a rock, tree, or building. Human-made materials, such as string, plastic, cloth, or paper, may also be used. Nests can be found in all types of habitat.

Nest building is driven by a biological urge known as the nesting instinct in birds and mammals. Generally each species has a distinctive style of nest. Nest complexity is roughly correlated with the level of parental care by adults. Nest building is considered a key adaptive advantage among birds, and they exhibit the most variation in their nests ranging from simple holes in the ground to elaborate communal nests hosting hundreds of individuals. Nests of prairie dogs and several social insects can host millions of individuals.

Kin selection is a process whereby natural selection favours a trait due to its positive effects on the reproductive success of an organism's relatives, even when at a cost to the organism's own survival and reproduction. Kin selection can lead to the evolution of altruistic behaviour. It is related to inclusive fitness, which combines the number of offspring produced with the number an individual can ensure the production of by supporting others (weighted by the relatedness between individuals). A broader definition of kin selection includes selection acting on interactions between individuals who share a gene of interest even if the gene is not shared due to common ancestry.

Charles Darwin discussed the concept of kin selection in his 1859 book, On the Origin of Species, where he reflected on the puzzle of sterile social insects, such as honey bees, which leave reproduction to their mothers, arguing that a selection benefit to related organisms (the same "stock") would allow the evolution of a trait that confers the benefit but destroys an individual at the same time. J.B.S. Haldane in 1955 briefly alluded to the principle in limited circumstances (Haldane famously joked that he would willingly die for two brothers or eight cousins), and R.A. Fisher mentioned a similar principle even more briefly in 1930. However, it was not until 1964 that W.D. Hamilton generalised the concept and developed it mathematically (resulting in Hamilton's rule) that it began to be widely accepted. The mathematical treatment was made more elegant in 1970 due to advances made by George R. Price. The term "kin selection" was first used by John Maynard Smith in 1964.