Semicircular canals in the context of Periosteum

The vestibular system, in vertebrates, is a sensory system that creates the sense of balance and spatial orientation for the function of coordinating movement with balance. Together with the cochlea, a part of the auditory system, it constitutes the labyrinth of the inner ear in most mammals.

As movements consist of rotations and translations, the vestibular system comprises two components: the semicircular canals, which indicate rotational movements; and the otoliths, which indicate linear accelerations. The vestibular system sends signals primarily to the neural structures that control eye movement; these provide the anatomical basis of the vestibulo-ocular reflex, which is required for clear vision. Signals are also sent to the muscles that keep an animal upright and in general control posture; these provide the anatomical means required to enable an animal to maintain its desired position in space.

The cochlear nerve (also auditory nerve or acoustic nerve) is one of two parts of the vestibulocochlear nerve, a cranial nerve present in amniotes, the other part being the vestibular nerve. The cochlear nerve carries auditory sensory information from the cochlea of the inner ear directly to the brain. The other portion of the vestibulocochlear nerve is the vestibular nerve, which carries spatial orientation information to the brain from the semicircular canals, also known as semicircular ducts.

The saccule (Latin: sacculus) is a bed of sensory cells in the inner ear that detects linear acceleration and head tilting in the vertical plane, and converts these vibrations into electrical impulses to be interpreted by the brain. When the head accelerates vertically, the sensory cells of the saccule are moved due to a combination of inertia and gravity. In response, the neurons connected to the saccule transmit electrical impulses that represent this movement to the brain. These impulses travel along the vestibulocochlear nerve (CNVIII) to the vestibular nuclei in the brainstem.

The vestibular system is important for balance, or equilibrium. It includes the saccule, utricle, and the three semicircular canals. The vestibule is the name of the fluid-filled, membranous duct that contains these organs of balance and is in turn encased in the temporal bone of the skull as a part of the inner ear.