Rauisuchus in the context of Santa María Formation


Rauisuchus in the context of Santa María Formation

⭐ Core Definition: Rauisuchus

Rauisuchus (meaning "Wilhelm Rau's crocodile") is a genus of extinct archosaurs which lived in what is now the Geopark of Paleorrota (Santa María Formation), Brazil, during the Late Triassic period (235–228 million years ago). It contains one species, R. tiradentes.

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Rauisuchus in the context of Phytosaur

Phytosaurs (Φυτόσαυροι in Greek, meaning 'plant lizard') are an extinct group of large, mostly semiaquatic Late Triassic archosauriform or basal archosaurian reptiles. Phytosaurs belong to the order Phytosauria and are sometimes referred to as parasuchians. Phytosauria, Parasuchia, Parasuchidae, and Phytosauridae have often been considered equivalent groupings containing the same species. Some recent studies have offered a more nuanced approach, defining Parasuchidae and Phytosauridae as nested clades within Phytosauria as a whole. The clade Phytosauria was defined by Paul Sereno in 2005 as Rutiodon carolinensis and all taxa more closely related to it than to Aetosaurus ferratus, Rauisuchus tiradentes, Prestosuchus chiniquensis, Ornithosuchus woodwardi, or Crocodylus niloticus (the Nile crocodile). Phytosaurs were long-snouted and heavily armoured, bearing a remarkable resemblance to modern crocodilians in size, appearance, and lifestyle, as an example of convergence or parallel evolution.

The name phytosaur means 'plant lizard', as the first fossils of phytosaurs were mistakenly thought to belong to plant-eaters.

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Rauisuchus in the context of Rauisuchian

"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large (often 4 to 6 metres (13 to 20 ft)), carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, phytosaurs (crocodile-like carnivores), aetosaurs (armored herbivores), and crocodylomorphs (lightly-built crocodilian ancestors).

However, more recent studies on archosaur evolution have upended this idea based on phylogenetic analyses and cladistics, a modern approach to taxonomy based on clades (nested monophyletic groups of common ancestry). Since the early 2010s, archosaur classification schemes have stabilized on a system where Rauisuchia is rendered an evolutionary grade, or even a wastebin taxon. Crocodylomorphs most likely originated from a rauisuchian ancestor based on a myriad of shared traits, and some "rauisuchians" (such as Postosuchus and Rauisuchus) appear to be more closely related to crocodylomorphs than to other "rauisuchians" (such as Prestosuchus and Saurosuchus).

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Rauisuchus in the context of Rauisuchidae

Rauisuchidae is a group of large (up to 6 metres (20 ft)) predatory Triassic archosaurs. Some disagreement exists over which genera should be included in the Rauisuchidae and which should be in the related Prestosuchidae and Poposauridae, and indeed whether these should even be thought of as separate valid families. Rauisuchidae in the modern sense was defined by Sterling Nesbitt in 2011 as the most inclusive clade containing Rauisuchus tiradentes, but not Prestosuchus chiniquensis, Poposaurus gracilis, or Crocodylus niloticus (the Nile crocodile). In this modern sense, rauisuchids are recovered as members of the clade Loricata, being the sister taxon of Crocodylomorpha (the group including living crocodilians), and being more derived than taxa such as Prestosuchus and Batrachotomus. Rauisuchids occurred throughout much of the Triassic, and may have first occurred in the Early Triassic if some archosaurian taxa such as Scythosuchus and Tsylmosuchus are considered to be within the family.

An early cladistic analysis of crocodylotarsan (pseudosuchian) archosaurs included Lotosaurus, Fasolasuchus, Rauisuchus, and "the Kupferzell rauisuchid" (later called Batrachotomus) within the Rauisuchidae. However, a later study found that Batrachotomus was a more basal pseudosuchian only slightly more "advanced" than Prestosuchus. In addition, the toothless Lotosaurus has been found to be more closely related to the Ctenosauriscidae, a clade of poposauroids with sails on their backs.

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